The aperture is an opening in certain kinds of mollusc shells: it is the main opening of the shell, where the head-foot part of the body of the animal emerges for locomotion, feeding, etc.
The word is not used to describe bivalve shells, where a natural opening between the two shell valves in the closed position is usually called a gape.
Scaphopod shells are tubular, and thus they have two openings: a main anterior aperture and a smaller posterior aperture.
As well as the aperture, some gastropod shells have additional openings in their shells for respiration; this is the case in some Fissurellidae (keyhole limpets) where the central smaller opening at the apex of the shell is called an orifice, and in the Haliotidae (abalones) where the row of respiratory openings in the shell are also called orifices.
The aperture of many snail shells is more or less round, rounded, elliptical or oval. This shape usually corresponds roughly to the cross-section of the body whorl of the shell.
The aperture of a snail shell can have many other forms: semicircular, trilobate or auriculate. In some gastropods, the aperture is narrowed by protruding plaits, which help make the soft parts of the animal less vulnerable to predation.
The growth of the shell is provided for by non-continuous addition of minute layers to the aperture margin (also called peristome) from the mantle border, the principal agent in the secretion of the shell.
The margin of the aperture is sometimes continuous or entire (Epitonium), or becomes continuous in the adult (Caracolus); very frequently it is interrupted, the left side of the aperture being formed only by the body whorl. When the aperture is called holostomatous, this means that the aperture is rounded or entire, uninterrupted by the siphonal canal, notch, or by any other extension
For convenience of reference, the margin of a gastropod aperture is divided into three areas:
The aperture is descending or deflected, when it does not follow the spiral of the shell, but turns downwards (such as in Helix). Sometimes it departs from contact with the preceding whorl (as frequently in Cylindrella.
The shape of the aperture in a gastropod shell can be:
Conus betulinus has a long and narrow aperture.
Indrella ampulla has an oval aperture.
Valvata sincera has a rounded aperture.
Theodoxus danubialis has a semicircular aperture.
Some shells, like this Patella vulgata, have no aperture.
The shells of juveniles in some species (especially some families of land snails) have a simple aperture with a sharp edge, but after reaching adult size the aperture of the shell finally acquires adult characters, consisting of a thickened, reflected, inflected or lipped edge, which is sometimes more or less contracted by inflected calcareous projections known as teeth. These teeth may be outer lip teeth, columellar teeth or parietal teeth.
Folds or plicae are named by their position in the aperture, as follows. The numbers refer to those in the diagram shown opposite:
Folds or plicae
Lamellae are named as follows: