Kota Formation

Summary

The Kota Formation is a geological formation in India. The age of the Kota Formation is uncertain; it is commonly considered to date to the Early Jurassic, but some studies have suggested it may extend into the Middle Jurassic or even later. It conformably overlies the Lower Jurassic Upper Dharmaram Formation and is unconformably overlain by the Lower Cretaceous Gangapur Formation. It is split into a Lower Member and Upper Member. The Lower Member is approximately 100 m thick while the Upper Member is 490 m thick. Both subunits primarily consist of mudstone and sandstone, but near the base of the upper unit there is a 20-30 metre thick succession of limestone deposited in a freshwater setting.[1]

Kota Formation
Stratigraphic range: Jurassic
TypeGeological formation
Sub-unitsLower & Upper members
UnderliesUnconformity with the Gangapur Formation
OverliesDharmaram Formation
Thickness550–600 m (1,800–1,970 ft)
Lithology
PrimaryMudstone, sandstone
OtherLimestone
Location
Coordinates18°54′N 80°00′E / 18.9°N 80.0°E / 18.9; 80.0
Approximate paleocoordinates31°36′S 31°54′E / 31.6°S 31.9°E / -31.6; 31.9
RegionTelangana
Country India
ExtentPranhita-Godavari Basin
Type section
Named forKota Village
Kota Formation is located in India
Kota Formation
Kota Formation (India)

Stratigraphy edit

The lower boundary of the Kota Formation is made of pebbly sandstone, covering the topmost clay seen in the Dharmaram Formation.[2] The Kota Formation has been traditionally divided into 2 main members, the Lower and Upper members, yet more recent work have redivided it into 3.[3] The Lower member can be seen at locations such as Adamilli, Kamavarapukota and Sudikonda, being made of sandstones, with clay clasts, with greater or lower stratification.[4] The Middle Member is well developed along the Continental Gondwana basin, specially towards the northwestern part, and is made of medium to fine white sandstone with clay and concretionary limestone, suggesting the development of paleosols associated with alluvial floodplains.[3] The last member is mostly made of broad sandstone sheets with large clay casts associated with fluvial channels, and has an extension that can be easuly seen on several continuous kilometers.[4][5] The Uppermost section of the unit is mostly made of limestones and is overlain on an angular unconformity by the Gangapur Formation.[3]

Age edit

The age of the Kota Formation is controversial. There are no magmatic rocks or volcanic ash beds associated with the Kota Formation, which means that its age cannot be determined directly through radiometric dating.[6][7] The maximum age of the Kota Formation is constrained by the underlying Upper Dharmaram Formation, which is Early Jurassic, probably Hettangian or Sinemurian, in age.[8][7] Various researchers have attempted to date the Kota Formation using biostratigraphy. Krishnan (1968), Jain (1973), and Yadagiri and Prasad (1977) favored an Early Jurassic age based on the fish fauna. Govindan (1975) suggested a Middle Jurassic age based on ostracods. In 2006, Bandyopadhyay and Sengupta argued that the fish fauna suggested a Toarcian age for the Upper Kota Formation, possibly extending into the Aalenian, and in turn estimated the Lower Kota to be Sinemurian to Pliensbachian in age.[9] Guntupalli V. R. Prasad, along with various coauthors, has argued for a younger age. In 2001, Vijaya and Prasad proposed based on palynological evidence that the Kota Formation was deposited between the Callovian age of the Middle Jurassic and the Barremian age of the Early Cretaceous.[10] In 2002, Prasad and Manhas argued that the mammal genus Dyskritodon, known only from the Kota Formation and the Early Cretaceous of Morocco, provides evidence for a young age for the Kota Formation.[11] In 2020, Prasad and Parmar argued that the similarity of the dinosaur fauna of the Kota Formation to that of the Middle Jurassic of the United Kingdom supported a Middle Jurassic age for the Kota Formation.[12]

Paleoenvironment edit

 
The Kota Formation hosted lacustrine and wetland settings, this last one being of carbonate type, having modern analoges such as the Tablas de Daimiel in Spain

The Kota Formation represents mostly a Continental succession related to a continental rift basin, the Pranhita-Godavari Gondwana Basin of peninsular India.[13] The associated facies of sandstone and limestones are likely related to playa-type lake, with nearby fluvial currents, part of low gradient hanging wall alluvial fans, being deposited on it´s margin. There have been records of freshwater lue green algal stromatolites and oncolites, suggested to be deposited on low energy and low bathymetry lacustrine settings.[14] More recent works have proven the basin hosted in the Early Jurassic a freshwater carbonate wetland marked by the presence of limestones.[1] The environmental model proposed include a depositional cycle marked by several facies types, A for the sublittoral zones of shallow water bodies, followed by palustrine environments, including surfaces with abundance of influence of both plants and animals, specially rhizobrecciation indicating active colonization of the margins by plants, having a similar deposition to the modern Las Tablas de Daimiel wetlands.[1] Associated with the lacustrine facies have recovered microbial bioherms and lacustrine spring mounds, shallow ephemeral ponds with carbonated mud and Phyllopods, pedogenic calcrete under arid seasons and short-lived distributary channels.[13] The depositional setting may have been partially sheltered from the input of siliciclastic materials, except on flooding seasons. Microbial biomats likely developed on shallow waters, while rhizoliths increased it´s presence of abandoned channel fills and pedogenic facies indicate drought seasons.[13]

Fossil content edit

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Ostracoda edit

Ostracods
Genus Species Location Stratigraphic position Material Notes Images

Clinocypris[15]

Clinocypris sp.

  • Daroghapalli
  • Upper Member

Six carapaces

A freshwater ostracodan of the Family Pontocyprididae.

Cypredea[15]

Cypredea sp.

  • Daroghapalli
  • Upper Member

Fourteen incomplete carapaces

A freshwater ostracodan of the Family Palaeocytheridae.

Darwinula[16][15]

Darwinula cf.sarytirmenensis

  • Aklapalli
  • Daroghapalli
  • Kanchelli
  • Metpalli
  • Potepalli
  • Lower Member
  • Upper Member

More than 200 carapaces and valves

A freshwater ostracodan of the Family Darwinulidae. The most dominant genus locally and the main indicator of both fluvial and lacustrine settings

Darwinula kingi

  • Aklapalli
  • Daroghapalli
  • Kanchelli
  • Metpalli
  • Potepalli
  • Lower Member
  • Upper Member

Around 120 carapaces and valves

A freshwater ostracodan of the Family Darwinulidae.

Darwinula spp.

  • Daroghapalli
  • Lower Member
  • Upper Member

Nineteen Carapaces

A freshwater ostracodan of the Family Darwinulidae.

Eucandona[15]

Eucandona sp.

  • Daroghapalli
  • Upper Member

Eight incomplete carapaces

A freshwater ostracodan of the Family Candoninae.

Limnocythere[16]

Limnocythere spp.

  • Daroghapalli
  • Lower Member

Three complete carapaces

A freshwater ostracodan of the family Limnocytheridae.

Stenocypris[15]

?Stenocypris sp.

  • Daroghapalli
  • Upper Member

Single incomplete carapace

A freshwater ostracodan of the family Cyprididae.

Timiriasevia[16]

Timiriasevia digitalis

  • Daroghapalli
  • Lower Member

Twenty complete carapaces and thirty-six partly broken carapaces.

A freshwater ostracodan of the family Limnocytheridae.

Phyllopoda edit

Phyllopods
Genus Species Location Stratigraphic position Material Notes Images

Estheriina[17][18]

Estheriina alibadadensis

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Estheriininae. The most abundant Estheriid in the region and the key element of the Estheriina biozone

Estheriina indijurassica

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Estheriininae.

Estheriina bullata

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Estheriininae.

Estheriina pranhitaensis

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Estheriininae.

Lioestheria[17][18]

Lioestheria kotaensis

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Lioestheriidae. The second key element of the Estheriina biozone

 

Lioestheria crustabundis[17]

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Lioestheriidae.

Lioestheria ssp.[17]

  • Kota limestone ridge
  • Lower Member

Valves

A freshwater clam shrimp of the family Lioestheriidae.

Paleolimnadia[17]

Paleolimnadia spp.

  • Kota limestone ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Estheriininae.

Pseudeasmussiata[17]

Pseudeasmussiata andhrapradeshia

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Lioestheriidae.

Insecta edit

Insects
Genus Species Location Stratigraphic position Material Notes Images

Archimesoblatta[19]

A. shiva

Tasch outcrop K-2 bed 8

  • Upper Member

Right forewing tegmen

A mesoblattinid cockroach

Coleopteron[20]

C. sp.

Kota limestone ridge

  • Lower Member

Isolated wings

An Indeterminate Coleopteran.

Kotaphialtites[21]

K. frankmortoni

Sirpur Taluka, Tasch's K1 outcrop bed 2(A)

  • Upper Member

MCZ 11909, Isolated wing

An ephialtitid hymenopteran.

Protogryllus[22]

P. lakshmi

Kota Formation outcrop K-2

  • Upper Member

MCZ 3046, Isolated wing

A protogryllid cricket

Taschigatra[23]

T. bharataja

Sirpur Taluka, Tasch's K1 outcrop bed 3(A)

  • Upper Member

No.2013(3013), part and counterpart of well preserved wing

A rhagionid dipteran.

T. tulyabhijana

Sirpur Taluka, Tasch's K1 outcrop bed 2(A)

  • Upper Member

No. 5034, well preserved wing

A rhagionid dipteran.

Xyelula[21]

X. alexandri

Kota Formation outcrop K-2

  • Upper Member

MCZ 11831, well preserved wing

A sepulcid hymenopteran.

Fish edit

Fish
Taxon Species Location Stratigraphic position Material Notes Images

Indocoelacanthus[24]

Indocoelacanthus robustus

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
  • ISI P. 39. Impression of the caudal fin
  • ISI P. 40. Nearly complete head and trunk squamation
  • ISI P. 41-45. Fragments of fin-rays
  • ISI P. 46. Neural arches, isolated scales and girdle elements

A robust freshwater coelacanth of the family Latimeriidae. Represents the largest member of the local freshwater fauna, measuring up to 70 cm.[24]

Lepidotes[25][26][27]

Lepidotes deccanensis

  • Kota limestone ridge
  • Paikasigudem village
  • Lower Member
  • Upper Member
  • Complete Specimen
  • Isolated Remains

A freshwater neopterygian of the family Lepisosteiformes.

 

Lepidotes spp.

  • Kota limestone ridge
  • Paikasigudem village
  • Lower Member
  • Upper Member

Isolated remains

A freshwater neopterygian of the family Lepisosteiformes.

Lonchidion[28][25]

Lonchidion indicus

  • Paikasigudem village
  • Yamanpalli bonebed
  • Upper Member
  • GSI.TI.2, tooth
  • Isolated Teeth

A freshwater elasmobranch of the family Lonchidiidae.

Paradapedium[25][29]

Paradapedium egertoni

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Near Ankisha
  • Lower Member
  • Upper Member
  • BMNHP12146, Partial specimen
  • BMNHP12147, Almost complete specimen
  • BMNHP12148, Partial specimen
  • ISIP.32 Nearly complete specimen
  • ISIP.33 Nearly complete specimen
  • ISIP.34 Nearly complete specimen
  • ISIP.35 specimen lacking tail and head

A freshwater neopterygian of the family Dapediidae.

Pholidophorus[30]

Pholidophorus kingi

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Paikasigudem village
  • Lower Member
  • Upper Member
  • Several complete and incomplete specimens

A freshwater neopterygian of the family Pholidophoridae.

Pholidophorus indicus

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Paikasigudem village
  • Lower Member
  • Upper Member
  • Several complete and incomplete specimens

A freshwater neopterygian of the family Pholidophoridae.

Polyacrodus[25][31]

Polyacrodus? sp.

  • Paikasigudem village
  • Upper Member
  • Tooth

A freshwater elasmobranch of the family Polyacrodontidae.

Pycnodontidae[28]

Indeterminate

  • Paikasigudem village
  • Upper Member
  • About 60 specimens of Pharyngeal Teeth, including GSITL3-6

A freshwater neopterygian of the family Pycnodontidae, originally classified as Perciformes, yet suggested to be very similar to the Cretaceous pycnodont Stephanodus.

Semionotiformes[25]

Indeterminate

  • Paikasigudem village
  • Upper Member
  • Teeth

A freshwater neopterygian of the family Semionotiformes.

Tetragonolepis[25][29]

Tetragonolepis oldhami

  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
  • GSI2145, an impression including most of the body
  • ISP36, fragment of trunk and skull
  • ISP37, nearly complete specimen

A freshwater neopterygian of the family Dapediidae.

 

Amphibia edit

Amphibians
Taxon Species Location Stratigraphic position Material Notes Images

Anura[28]

Indeterminate

  • Paikasigudem village
  • Upper Member
  • GSI.TL.19, Mandible
  • GSI.TL.20, right illium

Indeterminate frog remains, originally referred to Pelobatidae due to be compared with younger Creteaceous Indian frog material

Caudata[28]

Indeterminate

  • Paikasigudem village
  • Upper Member
  • GSI.TI.10, mandible with one teeth
  • GSI.TI.16, fragment of mandible
  • GSI.TI.17, madible with 4 teeth

Indeterminate caudatan remains, originally referred to Sirenidae due to be compared with younger Creteaceous sirenid material

Mammaliaforms edit

Mammaliaforms
Taxon Species Location Stratigraphic position Material Notes Images

Australosphenida[32]

Indeterminate

Paikasigudem village

  • Upper Member

Isolated lower molar

A mammal of the group Australosphenida, resembling the south american genus Asfaltomylos

Dyskritodon?[11]

Dyskritodon? indicus

Paikasigudem village

  • Upper Member

VPL/JU/KM/13, lower left molar

A dubious mammal of the group Eutriconodonta. This Genus is known from the Early Cretaceous of Morocco, what has been used to suggest a minimum Berrasian age for the Upper Kota Formation

Gondtherium[31]

Gondtherium dattai

Paikasigudem village

  • Upper Member

VPL/JU/KM 12 right lower molar

A mammal of the family Docodontidae

Indotherium[33]

Indotherium pranhitai

5 km west of Yamanapalli

  • Lower Member

GSI20795, right upper molar

A mammaliform of the family Morganucodontidae. Includes the informally named "Indozostrodon simpsoni".[34]

Indobaatar[35]

Indobaatar zofiae

Paikasigudem village

  • Upper Member

VPL/JU/KM/20, a left upper premolar

A mammal described as an eobaatarid multituberculate, but this interpretation has been challenged.[36]

Kotatherium[37]

Kotatherium haldanei

5 km west of Yamanapalli

  • Lower Member

GSI19634, right upper molar

A mammaliform of the family Kuehneotheriidae

Nakunodon[38]

Nakunodon paikasiensis

Paikasigudem village

  • Upper Member

GSI.SR/PAL/12, right upper molar

A mammal of the family Amphidontidae

Paikasigudodon[11][39]

Paikasigudodon yadagirii

Paikasigudem village

  • Upper Member

VPL/JU/KM/10, right upper molar

A mammaliaform of the family Morganucodontidae, originally known as "Kotatherium yadagirii"

Trishulotherium[33]

Trishulotherium kotaensis

Paikasigudem village

  • Upper Member

GSISR/PAL/10, left lower molar

A mammal of the order Symmetrodonta

Lepidosauromorpha edit

Lepidosauromorphs
Taxon Species Location Stratigraphic position Material Notes Images

Bharatagama[40]

Bharatagama rebbanensis

Paikasigudem village

  • Upper Member
  • VPL/JU/KR 66, dentary
  • VPL/JU/KR 88, anterior region of a left maxilla
  • VPL/JU/KR 91, left maxilla
  • VPL/JU/KR 90-92, posterior ends of right maxillae
  • VPL/JU/KR 67-80-103, anterior symphysial region
  • JU/KR 79, 98, 100, dentaries mid-region
  • VPL/JU/KR 69, 81-84, 87, dentary with hatchling dentition
  • VPL/JU/KR 68, 85-86, dentary
  • VPL/JU/KR 70, 71, 76,78, 93, 94, 97, 104, 105, dentary

A lepidosauromorph originally described as an Iguanian lizard. May actually be a sphenodontian rather than a lizard.[41]

Godavarisaurus[42]

Godavarisaurus lateefi

  • Gorlapalli village
  • Paikasigudem village
  • Upper Member
  • VP4Ju/KR47, a partial right dentary
  • VP4Ju/KR44, 51, 52, right maxilla
  • VP4Ju/KR40, 41, 43, 49, 50, left maxilla
  • VP4Ju/KR37, 48, 65, right dentary
  • VP4Ju/KR39, fragment of left dentary
  • VP4Ju/KR46, left palatine

A small sphenodontian, with a skull estimated to measure less than 20 mm

Paikasisaurus[28][40]

Paikasisaurus indicus

Paikasigudem village

  • Upper Member
  • GSI.TI.14, left dentary with teeth
  • GSI.TI.15, left dentary with one tooth

An indeterminate and dubious lepidosauromorph, originally suggested to be a varanoid lizard

Rebbanasaurus[42]

Rebbanasaurus jaini

Paikasigudem village

  • Upper Member
  • About 50 specimens, including premaxillae, maxillae, dentaries, and a palatine

A small sphenodont

Squamata?[28][40]

Indeterminate

Paikasigudem village

  • Upper Member
  • VPL/JU/KR 62, fragmentary right maxilla
  • VPL/JU/KR 61, partial maxilla

Distinct from Bharatagama rebbanensis; may include material formerly assigned to the dubious Kota squamate Paikasisaurus indicus.[a]

Sphenodontidae[28][42]

Indeterminate

Paikasigudem village

  • Upper Member
  • GSI.TI.8-12, 17, maxilla

Indeterminate Sphenodontidae remains

Testudinata edit

Turtles
Taxon Species Location Stratigraphic position Material Notes Images

Indochelys[43][44]

Indochelys spatulata

  • 3 km NNE of Kistapur village, Kota
  • Near Kota village, north of Sironcha
  • Upper Member
  • GSI 20380, a partial shell
  • ISI R176, a partial shell
  • ISI R177, the anterior half of a shell

Mesochelydian stem-turtle, suggested to be related with Condorchelys

Testudines[45]

Indeterminate

1 km south of Bodepalli

  • Lower Member

Carapace fragments

Indeterminate turtle remains

Crocodylomorpha edit

Atoposauridae crocodiles are known from the unit, yet is not clear from what locality.[7]

Crocodylomorpha
Taxon Species Location Stratigraphic position Material Notes Images

Crocodylomorpha[46]

Indeterminate

1 km south of Bodepalli

Lower Member

Maxillae, dentaries, teeth

Indeterminate crocodylomorph remains, previously mixed with thyreophoran material and part of the chimaeric "Andhrasaurus"

Teleosauridae?[47]

Indeterminate

Kota limestone ridge

  • Lower Member

Dermal scutes, with a femur and some fragments of other bones

Indeterminate material referred to crocodylomorphs similar to Teleosaurus

Pterosauria edit

Pterosaurs
Taxon Species Location Stratigraphic position Material Notes Images

Campylognathoides[48]

Campylognathoides indicus

Kota limestone ridge

  • Lower Member
  • ISI R38, holotype, fragment of skull and upper jaw

The holotype of Campylognathoides indicus, a pair of premaxillae, may represent a fish rather than a pterosaur.[49]

Pterosauria[48][50]

Indeterminate

  • Boraigudem
  • Chitur
  • Lower Member
  • GSI17868, radius and ulna, wing metacarpal, first wing-phalanx and a clawed toe
  • ISIR49. fragment displaying dissociated bones from post-cranial region

Indeterminate pterosaur remains

Dinosaurs edit

Dinosaurs
Genus Species Location Stratigraphic position Material Notes Images

"Andhrasaurus"[45][51]

"Andhrasaurus indicus"

1 km south of Bodepalli

Lower Member

Sacral vertebra, vertebral centra, dorsal vertebrae, caudal vertebrae, parts of scapula and ilium, osteoderms

A chimaera of thyreophoran postcranial material and Crocodrylomorph skull pieces. The armor was later suggested to be Ankylosauria indet.[46] And other latter works pointed out it likely belongs to an indeterminate basal thyreophoran.[52]

Barapasaurus[53]

Barapasaurus tagorei

  • North of Krishnapur
  • Pochampalli village
  • Lower Member
  • Roughly 300 bones, representing the disarticulated remains of at least six individuals

A sauropod dinosaur, either a Eusauropoda or more likely a Gravisauria. Represents the best-known Early Jurassic sauropod

 

"Dandakosaurus"[54]

"Dandakosaurus indicus"

Yamanpalli bonebed

Lower Member

  • Lateral tooth, proximal ischium
  • Dorsal vertebra, proximal caudal vertebra (Likely of Kotasaurus)

A chimaera of large theropod bones, including and ischium and tooth, probably belonging to a carnosaur, and sauropod bones (two Kotasaurus vertebrae)

 

Dromaeosauridae?[12]

Indeterminate

Paikasigudem village

  • Upper Member

Isolated Teeth

Five distinct morphotypes have been identified, mostly resemble coelurosaurs or dromaeosauroids.[12]

Hypsilophodontidae[55]

Indeterminate

Gorlapalli Village

  • Upper Member
  • Single Tooth

Identified originally as a member of Hypsilophodontidae, probably represents a tooth of a basal neornithischian.[12]

Kotasaurus[56]

Kotasaurus yamanpalliensis

Yamanpalli bonebed

  • Lower Member

Disarticulated remains of at least 12 individuals[56]

A basal sauropod

 

Ornithischia[46][12]

Indeterminate

  • 1 km south of Bodepalli
  • Paikasigudem village
  • Lower Member
  • Upper Member
  • End of left humerus
  • Isolated Teeth

Indeterminate ornithischian material. Among the teeth, at least five distinct morphotypes have been identified.

Richardoestesia?[12]

Richardoestesia? spp.

Paikasigudem village

  • Upper Member

Isolated Teeth

Teeth similar to those of the problematic taxon Richardoestesia, of supposed coelurosaur affinities

Thyreophora[12][57]

Indeterminate

Paikasigudem village

  • Upper Member

Scute and associated fragmentary limb bones

Indeterminate scelidosaurid material

Megaflora edit

Genus Species Location Stratigraphic position Material Notes Images

Agathoxylon[58]

Agathoxylon kotaense

Near Kota village

  • Lower Member

Fossil wood

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

Agathoxylon chandrapurensis[59]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

Agathoxylon santacruzense[60]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

Agathoxylon pranhitaensis[58][61]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

Agathoxylon santalense[58]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

Agathoxylon spp.[58]

  • Bodepalli
  • Near Kota village
  • Paikasigudem village
  • Lower Member
  • Upper Member

Fossil wood

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

Araucarites[61][50]

Araucarites minutus

  • Dronadula
  • Lower Member

Branched shoots

Affinities with Araucariaceae inside Pinales.

Araucarites sp.

  • Near Kota village
  • Lower Member

Branched shoots

Affinities with Araucariaceae inside Pinales.

Brachyphyllum[61][59]

Brachyphyllum expansum

  • Near Kota village
  • Lower Member

Branched shoots

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

Elatocladus[61][50]

Elatocladus conferta

Near Kota village

  • Lower Member

Branched shoots

Affinities with Cupressaceae inside Pinales.

Elatocladus tenerrimus[59]

Near Kota village

  • Lower Member

Branched shoots

Affinities with Cupressaceae inside Pinales.

Elatocladus jabalpurensis[59]

Near Kota village

  • Lower Member

Branched shoots

Affinities with Cupressaceae inside Pinales.

Elatocladus plana[59]

Near Kota village

  • Lower Member

Branched shoots

Affinities with Cupressaceae inside Pinales.

Elatocladus sp.[59]

Near Kota village

  • Lower Member

Branched shoots

Affinities with Cupressaceae inside Pinales.

Cladophlebis[50]

Cladophlebis denticulata

  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member

Isolated fronds

Affinities with Osmundaceae in the Osmundales.

 
Cladophlebis denticulata illustration

Cladophlebis indica[61][50]

  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member

Isolated fronds

Affinities with Osmundaceae in the Osmundales

Cladophlebis reversa[61][50]

  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member

Isolated fronds

Affinities with Osmundaceae in the Osmundales

Cladophlebis spp.[61][50]

  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member

Isolated fronds

Affinities with Osmundaceae in the Osmundales

Circoporoxylon[62]

Circoporoxylon kotaense

Near Kota village

  • Lower Member

Fossil wood

Affinities with Podocarpaceae inside Pinales.

Coniopteris[61][50]

Coniopteris hymenophylloides

  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member

Isolated fronds

Affinities with Polypodiales in the Polypodiidae. Common cosmopolitan Mesozoic fern genus. Recent research has reinterpreted it a stem group of the Polypodiales (closely related to the extant genera Dennstaedtia, Lindsaea, and Odontosoria)

 
Coniopteris specimen

Coniopteris sp.[61]

  • Near Kota village
  • Lower Member

Isolated fronds

Affinities with Polypodiales in the Polypodiidae.

Cupressinoxylon[58][61]

Cupressinoxylon kotaense

Near Kota village

  • Lower Member

Fossil wood

Affinities with Cupressaceae inside Pinales.

Dictyozamites[59]

Dictyozamites falcatus

  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

Dictyozamites kotaense[59]

  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

Equisetum[61]

Equisetum rajmahalensis

  • Chitur village
  • Near Kota village
  • Lower Member
  • Upper Member

Isolated Stems

Affinities with Equisetaceae inside Equisetales.

 
Extant Equisetum arvense specimen

Ginkgoites[61]

Ginkgoites lobata

  • Near Kota village
  • Lower Member

Leafs

Affinities with Ginkgoaceae inside Ginkgoopsida.

 
Ginkgoites reconstruction

Ginkgoxylon[63]

Ginkgoxylon dixii

Near Kota village

  • Lower Member

Fossil wood

Affinities with Ginkgoaceae inside Ginkgoopsida.

Hausmannia[61]

Hausmannia cf. buchii

Near Kota village

  • Lower Member

Isolated pinnae

Affinities with Dipteridaceae in the Polypodiales.

 
Hausmannia specimen

Otozamites[59]

Otozamites vemavarmensis

  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

 
Otozamites specimen

Pagiophyllum[61][50]

Pagiophyllum peregrinum

Near Kota village

  • Lower Member

Branched shoots

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

Pagiophyllum cf.peregrinum

Near Kota village

  • Lower Member

Branched shoots

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

Pagiophyllum spp.

Near Kota village

  • Lower Member

Branched shoots

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

Pachypteris[59]

Pachypteris indica

Near Kota village

  • Lower Member

Isolated pinnae

Affinities with Umkomasiaceae in the Pteridospermatophyta.

Planoxylon[59]

Planoxylon mahabalei

Near Kota village

  • Lower Member

Fossil wood

Affinities with Protopinaceae inside Pinales.

Podocarpoxylon[61]

Podocarpoxylon chandrapurensis

Near Kota village

  • Lower Member

Fossil wood

Affinities with Podocarpaceae inside Pinales.

Podocarpoxylon chiturensis[59]

Chitur village

  • Upper Member

Fossil wood

Affinities with Podocarpaceae inside Pinales.

Podocarpoxylon krauselii[61]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Podocarpaceae inside Pinales.

Podocarpoxylon rajmahalense[61]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Podocarpaceae inside Pinales.

Podocarpoxyion sewardii[59]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Podocarpaceae inside Pinales.

Podocarpoxylon sp.[61]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Podocarpaceae inside Pinales.

Podozamites[61]

Podozamites sp.

  • Near Kota village
  • Lower Member

Leaflets

Broad conifer leaves

Pseudoctenis[59]

Pseudoctenis cf. frngilis

  • Near Kota village
  • Lower Member

Leaflets

Affinities with Cycadales in the Cycadopsida.

Ptilophyllum[61]

Ptilophyllum fissum

  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

Ptilophyllum acutifolium

  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

Ptilophyllum cutchense[59]

  • Chitur village
  • Near Kota village
  • Lower Member
  • Upper Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

Ptilophyllum cf.sahnii[59]

  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

Ptilophyllum cf.institacallum[59]

  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

Ptilophyllum sp.

  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

Protaxodioxylon[64]

Protaxodioxylon sahnii

Chitur village

  • Upper Member

Fossil wood

Affinities with Cupressaceae inside Pinales.

Prototaxoxylon liassicum[65]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Cupressaceae inside Pinales.

Sphenopteris[61][59]

Sphenopteris kotaensis

Near Kota village

  • Lower Member

Isolated Fronds

Affinities with Dicksoniaceae in the Cyatheales.

Taxaceoxylon[61]

Taxaceoxylon sahnii

Near Kota village

  • Lower Member

Fossil wood

Affinities with Cupressaceae inside Pinales.

Taxaceoxylon biradarii[59]

Chitur village

  • Lower Member

Fossil wood

Affinities with Cupressaceae inside Pinales.

Taxaceoxylon antiquum[59]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Cupressaceae inside Pinales.

Taxaceoxylon sp.

Near Kota village

  • Upper Member

Fossil wood

Affinities with Cupressaceae inside Pinales.

Torreyites[59]

Torreyites constricte

  • Chitur village
  • Upper Member

Branched shoots

Affinities with Taxaceae in the Pinales.

 
Extant Torreya specimen

See also edit

Notes edit

  1. ^ It cannot be determined which of the two Kota squamate forms, if either, the holotype of Paikasisaurus indicus pertains to.[40]

References edit

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