The Latins (Latin: Latini), sometimes known as the Latians, were an Italic tribe which included the early inhabitants of the city of Rome. From about 1000 BC, the Latins inhabited the small region known to the Romans as Old Latium (in Latin Latium vetus), that is, the area between the river Tiber and the promontory of Mount Circeo 100 km (62 mi) southeast of Rome. Following the Roman expansion, the Latins spread into the Latium adiectum, inhabited by Osco-Umbrian peoples.
Their language, Latin, belonged to the Italic branch of Indo-European. Speakers of Italic languages are assumed to have migrated into the Italian Peninsula during the late Bronze Age (1200–900 BC). The material culture of the Latins, known as the Latial culture, was a distinctive subset of the proto-Villanovan culture that appeared in parts of the Italian peninsula in the first half of the 12th century BC. The Latins maintained close culturo-religious relations until they were definitively united politically under Rome in 338 BC, and for centuries beyond. These included common festivals and religious sanctuaries.
The rise of Rome as by far the most populous and powerful Latin state from c. 600 BC led to volatile relations with the other Latin states, which numbered about 14 in 500 BC. In the period of the Tarquin monarchy (c. 550–500 BC), Rome apparently acquired political hegemony over the other states. After the fall of the Roman monarchy around 500 BC, there appears to have been a century of military alliance between Rome and the other Latin states to confront the threat posed to all Latium by raiding by the surrounding Italic mountain tribes, especially the Volsci and Aequi. This system progressively broke down after roughly 390 BC, when Rome's aggressive expansionism led to conflict with other Latin states, both individually and collectively. In 341–338 BC, the Latin states jointly fought the Latin War against Rome in a final attempt to preserve their independence. The war resulted in 338 BC in a decisive Roman victory. The other Latin states were either annexed or permanently subjugated to Rome.
The name Latium has been suggested to derive from the Latin word latus ("wide, broad"), referring, by extension, to the plains of the region (in contrast to the mainly mountainous Italian peninsula). If this is true, then Latini originally meant "men of the plain".
The Latins belonged to a group of Indo-European-speaking (IE) tribes, conventionally known as the Italic tribes, that populated central and southern Italy during the Italian Iron Age, which began around 900 BC. The most widely accepted theory suggests that Latins and other proto-Italic tribes first entered Italy in the late Bronze Age proto-Villanovan culture, then part of the central European Urnfield culture system. In particular various authors, such as Marija Gimbutas, had noted important similarities between the proto-Villanovan culture, the South-German Urnfield culture of Bavaria-Upper Austria  and Middle-Danube Urnfield culture. According to David W. Anthony proto-Latins originated in today's eastern Hungary, kurganized around 3100 BC by the Yamna culture, while Kristian Kristiansen associated the proto-Villanovans with the Velatice-Baierdorf culture of Moravia and Austria. This is further confirmed by the fact that the subsequent Latial culture, Este culture and Villanovan culture, which introduced iron-working to the Italian peninsula, were so closely related to the Central European Urnfield culture (c. 1300–750 BC), and Hallstatt culture (which succeeded the Urnfield culture), that it is not possible to tell them apart in their earlier stages. Furthermore, the contemporary Canegrate culture of Northern Italy represented a typical western example of the western Hallstatt culture, whose diffusion most probably took place in a Celtic-speaking context.
Similarly, several authors have suggested that the Beaker culture of Central and Western Europe was a candidate for an early Indo-European culture, and more specifically, for an ancestral European branch of Indo-European dialects, termed "North-west Indo-European", ancestral to Celtic, Italic, Germanic and Balto-Slavic. All these groups were descended from Proto-Indo-European speakers from Yamna-culture, whose migrations in Central Europe probably split off Pre-Italic, Pre-Celtic and Pre-Germanic from Proto-Indo-European.
Leaving archaeology aside, the geographical distribution of the ancient languages of the peninsula may plausibly be explained by the immigration of successive waves of peoples with different languages, according to Cornell. On this model, it appears likely that the "West Italic" group (including the Latins) were the first wave, followed, and largely displaced by, the East Italic (Osco-Umbrian) group. This is deduced from the marginal locations of the surviving West Italic niches. Besides Latin, putative members of the West Italic group are Faliscan (now regarded as merely a Latin dialect), and perhaps Sicel, spoken in central Sicily. The West Italic languages were thus spoken in limited and isolated areas, whereas the "East Italic" group comprised the Oscan and Umbrian dialects spoken over much of central and southern Italy. The chronology of Indo-European immigration remains elusive, as does the relative chronology between the Italic IE languages and the non-IE languages of the peninsula, notably the Etruscan. Most scholars consider that Etruscan is a pre-IE survival, part of a Mediterranean linguistic substratum. Some authors believe that, before the spread of the Gaulish language in the plain of the river Po from c. 400 BC onwards, northern and central Italy were dominated by non-IE languages: Etruscan, which shared some similarities with the Raetic, the possibly non-IE Ligurian and the language of the undeciphered Novilara inscriptions from the region around Ancona on the Adriatic coast. Some scholars have earlier suggested that Etruscan language could have been introduced by later migrants. The ancient Greek historian Herodotus preserves the tradition that the Tyrrhenoi (Etruscans) originated in Lydia in Anatolia, but Lydians spoke an Indo-European language, completely different from the Etruscan language. Possible support for an eastern origin for Etruscan may be provided by two inscriptions in a language closely related to Etruscan found on the island of Lemnos in the northern Aegean sea (see Lemnian language), although several scholars believe that the Lemnian language might have arrived in the Aegean Sea during the Late Bronze Age, when Mycenaean rulers recruited groups of mercenaries from Sicily, Sardinia and various parts of the Italian peninsula. Language change can be explained by scenarios other than mass migration e.g. by small immigrant-elites who acquire political/cultural hegemony over the indigenous majority and convert the latter to their language, a process known in historical linguistics as "elite transfer"., but there is no archaeological evidence in favor of an eastern origin of the Etruscans or the arrival of small immigrant-elites who may have imposed their language. Genetic studies on Etruscan samples, both on mitochondrial and autosomal DNA, are also against an eastern origin and have supported a deep, local origin. A 2019 Stanford genetic study, which has analyzed the autosomal DNA of Iron Age samples from the areas around Rome, has concluded that Etruscans were similar to the Latins from Latium vetus. According to British archeologist Phil Perkins, "there are indications that the evidence of DNA can support the theory that Etruscan people are autochthonous in central Italy".
The oldest extant inscription in the Latin language is believed to be engraved on the Lapis Niger ("Black Stone") discovered in 1899 in the Roman Forum, dating from around 600 BC: in the mid-Roman kingdom, according to the traditional Roman chronology, but more likely close to its inception. Written in a primitive form of Archaic Latin, it indicates that the Romans remained Latin-speakers in the period when some historians have suggested that Rome had become "Etruscanised" in both language and culture. It also lends support to the existence of the Kings of Rome in this era, whom some historians regarded as mythical: the inscription contains the word recei, the word for "king" in the dative singular in archaic Latin - regi in classical Latin, or to the rex sacrorum, rather than the political king of Rome.
There is no archaeological evidence at present that Old Latium hosted permanent settlements during the Bronze Age. Some very small amounts of Apennine culture pottery shards have been found in Latium, most likely belonging to transient pastoralists engaged in transhumance. It thus appears that the Latins occupied Latium Vetus not earlier than around 1000 BC. Initially, the Latin immigrants into Latium were probably concentrated in the low hills that extend from the central Apennine range into the coastal plain (much of which was then marshy and malarial, and thus uninhabitable). A notable area of early settlement were the Alban Hills, a plateau about 20 km (13 mi) SE of Rome containing a number of extinct volcanoes and 5 lakes, of which the largest are lacus Nemorensis (Lake Nemi) and lacus Tusculensis (Lake Albano). These hills provided a defensible, well-watered base. Also the hills on the site of Rome, certainly the Palatine and possibly the Capitoline and the Quirinal, hosted permanent settlements at a very early stage.
The Latins appear to have become culturally differentiated from the surrounding Osco-Umbrian Italic tribes from c. 1000 BC onwards. From this time, the Latins exhibit the features of the Iron Age Villanovan culture found in Etruria and the Po valley. In contrast, the Osco-Umbrian tribes do not exhibit Villanovan features. The Latins thus shared the broadly same material culture as the Etruscans. The variant of Villanovan found in Latium is dubbed the Latial culture. The most distinctive feature of Latial culture were cinerary urns in the shape of miniature tuguria ("huts"). In Phase I of the Latium culture (c. 1000–900 BC) these hut-urns only appear in some burials, but they become standard in Phase II cremation burials (900–770 BC). They represent the typical single-roomed hovels of contemporary peasants, which were made from simple, readily available materials: wattle-and-daub walls and straw roofs supported by wooden posts. The huts remained the main form of Latin housing until about 650 BC. The most famous exemplar was the Casa Romuli ("Hut of Romulus") on the southern slope of the Palatine Hill, supposedly built by the legendary founder of Rome with his own hands and which reportedly survived until the time of emperor Augustus (ruled 30 BC - AD 14).
Around 650 BC began a period of urbanisation, with the establishment of political city-states in Latium. The most notable example is Rome itself, which was originally a group of separate settlements on the various hills. It appears that they coalesced into a single entity around 625 BC, when the first buildings were established on the site of the later Roman Forum.
According to the mainstream Kurgan hypothesis, the earliest Indo-Europeans were a nomadic steppe people, originating in the Eurasian steppes (southern Russia, northern Caucasus and central Asia). Their livelihood was based on horses and herding. In the historical era, the same socio-cultural lifestyle was maintained, in the same regions, by peoples descended from the Proto-Indo-Europeans (PIEs) known to the Greco-Romans as Scythians, Sarmatians and Alans, whose languages belonged to the Iranian branch of IE. On the basis of common steppe-nomadic features in the cultures of the various Indo-European peoples in the historical era, scholars have reconstructed elements of proto-Indo-European culture. Relics of such elements have been discerned in Roman and Latin customs. Examples include:
Despite their frequent internecine wars, the Latin city-states maintained close culturo-religious relations throughout their history. Their most important common tribal event was the four-day Latiar or Feriae Latinae ("Latin Festival"), held each winter on the sacred mons Albanus (Monte Cavo, Alban Hills, SE of Rome), an extinct volcano. The climax of the festival was a number of sacrifices to Jupiter Latiaris ("Jupiter of Latium"); the sacrificed meat was shared by the representatives of the Latin communities. These elaborate rituals, as did all Roman religious ceremonies, had to be performed with absolute precision and, if any procedural mistakes were made, had to be repeated from the start. The Latin Festival continued to be held long after all Latium Vetus was integrated into the Roman Republic after 338 BC (from then on, the Roman consuls presided over them) and into the Roman imperial era. The historian Livy, writing around AD 20, ascribed Rome's disastrous defeat by the Carthaginian general Hannibal at the Battle of Lake Trasimene in 217 BC to the impiety of the consul Gaius Flaminius, who, in his eagerness to join his army at its assembly-point of Arretium, omitted to attend the Latin Festival.
Latin cultural-religious events were also held at other common cult-centres e.g. the major common shrine to Diana at Aricia. This may be the sacred grove to Diana which a fragment of Cato's Origines recorded dedicated, probably c. 500 BC, by various Latin communities under the leadership of the dictator of Tusculum, Egerius Baebius. Cornell argues that the temple of Diana reportedly founded by the Roman king Servius Tullius on the Aventine hill at Rome was also a common Latin shrine, as it was built outside the pomerium or City boundary. There was also an important Latin cult-centre at Lavinium. Lavinium hosted the cult of the Penates, or Latin ancestor-gods. Cornell suggests that the "Sanctuary of the 13 altars" discovered in the 1960s at Lavinium was the site of the Penates cult. Since each of the altars differ in style and date, it has been suggested that each was erected by a separate Latin city-state.
Under the ever-growing influence of the Italiote Greeks, the Romans acquired their own national origin-legend sometime during the early Republican era (500-300 BC). It was centred on the figure of Aeneas, a supposed Trojan survivor of the destruction of Troy by the Achaean Greeks, as related in the poet Homer's epic the Iliad (composed c. 800 BC). The legend provided the Romans with a heroic "Homeric" pedigree, as well as a (spurious) ethnic distinctiveness from the other Latins. It also provided a rationale (as poetic revenge for the destruction of Troy) for Rome's hostilities against, and eventual subjugation of, the Greek cities of southern Italy, especially Taras (mod. Taranto) in the period ending 275 BC.
The figure of Aeneas as portrayed in the Iliad lent itself to his adoption as the Roman "Abraham": a mighty warrior of (minor) royal blood who personally slew 28 Achaeans in the war, he was twice saved from certain death by the gods, implying that he had a great destiny to fulfil. A passage in Homer's Iliad contains the prophecy that Aeneas and his descendants would one day rule the Trojans. Since the Trojans had been expelled from their own city, it was speculated that Aeneas and other Trojan survivors must have migrated elsewhere.
The legend is given its most vivid and detailed treatment in the Roman poet Virgil's epic, the Aeneid (published around AD 20). According to this, the Latin tribe's first king was Latinus, who gave his name to the tribe and founded the first capital of the Latins, Laurentum, whose exact location is uncertain. The Trojan hero Aeneas and his men fled by sea after the capture and sack of their city, Troy, by the Greeks in 1184 BC, according to one ancient calculation. After many adventures, Aeneas and his Trojan army landed on the coast of Latium near the mouth of the Tiber. Initially, King Latinus attempted to drive them out, but he was defeated in battle. Later, he accepted Aeneas as an ally and eventually allowed him to marry his daughter, Lavinia. Aeneas supposedly founded the city of Lavinium (Pratica di Mare, Pomezia), named after his wife, on the coast not far from Laurentum. It became the Latin capital after Latinus' death. Aeneas' son (by his previous Trojan wife, a daughter of king Priam of Troy), Ascanius, founded a new city, Alba Longa in the Alban Hills, which replaced Lavinium as capital city. Alba Longa supposedly remained the Latin capital for some 400 years under Aeneas' successors, the Latin kings of Alba, until his descendant (supposedly in direct line after 15 generations) Romulus founded Rome in 753 BC. Under a later king Tullus Hostilius (traditional reign-dates 673–642 BC), the Romans razed Alba Longa to the ground and resettled its inhabitants on the mons Caelius (Caelian Hill) in Rome.
There is controversy about how and when Aeneas and his Trojans were adopted as ethnic ancestors by the Romans. One theory is that the Romans appropriated the legend from the Etruscans. There is evidence that the Aeneas legend was well known among the Etruscans by 500 BC: excavations at the ancient Etruscan city of Veii discovered a series of statuettes portraying Aeneas fleeing Troy carrying his father on his back, as in the legend. Indeed, the Bulgarian linguist Vladimir Georgiev argued that the original Etruscans were in fact descendants of those Trojan refugees and that the Aeneas legend has a historical basis. Georgiev disputes the mainstream view that Etruscan was not Indo-European: he argues that Etruscan was closely related to the Indo-European Hittite and Lydian languages, on the basis of lexical and morphological correspondences.
Georgiev's thesis has received only minority support from other scholars. But even if Etruscan was not Indo-European, that does not necessarily invalidate a possible Trojan origin for the Etruscans, as the everyday language of Troy in this period remains uncertain. Excavations at Troy have yielded a single written document, a letter in Luwian. But as Luwian (which certainly is closely related to Hittite) was used as a kind of diplomatic lingua franca in Anatolia, it cannot be argued conclusively that Luwian was the everyday language of Troy. Possible support for a Trojan-Etruscan link may be found in the existence of the so-called Lemnian language, which was spoken on the Aegean island of Lemnos at least until 500 BC, and is apparently closely related to Etruscan. Lemnos is close to the shore of Troy, and may have been a colony of the Trojans and/or a major destination for Trojan refugees after the destruction of their city. Cornell points out that the Romans may have acquired the legend directly from the Italiote Greeks. The earliest Greek literary reference to Rome as a foundation of Aeneas dates to c. 400 BC. There is also much archaeological evidence of contacts between the cities of archaic Latium and the Greek world e.g. the archaic sanctuary of the Penates at Lavinium, which shows "heavy Greek influence in architectural design and religious ideology", according to Cornell.
But whatever the origin of the legend, it is clear that the Latins had no historical connection with Aeneas and none of their cities were founded by Trojan refugees. Furthermore, Cornell regards the city of Alba Longa itself as probably mythical. Early Latial-culture remains have been discovered on the shore of the Alban lake, but they indicate a series of small villages, not an urbanised city-state. In any case, traces of the earliest phase of Latial culture also occur at Rome at the same time (c. 1000 BC), so archaeology cannot be used to support the tradition that Rome was founded by people from Alba Longa. If Alba Longa did not exist, then nor did the "Alban kings", whose genealogy was almost certainly fabricated to "prove" Romulus' descent from Aeneas. The genealogy's dubious nature is shown by the fact that it ascribes the 14 Alban kings an average reign of 30 years' duration, an implausibly high figure. The false nature of the Aeneas-Romulus link is also demonstrated by the fact that, in some early versions of the tradition, Romulus is denoted as Aeneas' grandson, despite being chronologically separated from Aeneas by some 450 years.
Romulus himself was the subject of the famous legend of the suckling she-wolf (lupa) that kept Romulus and his twin Remus alive in a cave on the Palatine Hill (the Lupercal) after they had been thrown into the river Tiber on the orders of their wicked uncle, Amulius. The latter had usurped the throne of Alba from the twins' grandfather, king Numitor, and then confined their mother, Rhea Silvia, to the Vestal convent. They were washed ashore by the river, and after a few days with the wolf, were rescued by shepherds.
Mainstream scholarly opinion regards Romulus as an entirely mythical character, and the legend fictitious. On this view, Romulus was a name fabricated to provide Rome with an eponymous founding hero, a common feature of classical foundation-myths; it is possible that Romulus was named after Rome instead of vice versa. The name contains the Latin diminutive -ulus, so it means simply "Roman" or "little Roman". It has been suggested that the name "Roma" was of Etruscan origin, or that it was derived from the Latin word ruma ("teat"), presumably because the shape of the Palatine Hill and/or Capitoline Hill resembled a woman's breasts. If Romulus was named after the city, it is more likely that he was historical. Nevertheless, Cornell argues that "Romulus probably never existed... His biography is a complex mixture of legend and folk-tale, interspersed with antiquarian speculation and political propaganda".
In contrast, Andrea Carandini, an archaeologist who has spent most of his career excavating central Rome, advanced the theory that Romulus was a historical figure who indeed founded the city in c. 753 BC, as related by the ancient chroniclers, by ploughing a symbolic sacred furrow to define the city's boundary. But Carandini's views have received scant support among fellow scholars.
In contrast to the legend of Aeneas, which was clearly imported into the Latin world from an extraneous culture, it appears that the Romulus legend of the suckling she-wolf is a genuine indigenous Latin myth.
The traditional number of Latin communities for the purposes of the joint religious festivals is given as 30 in the sources. The same number is reported, probably erroneously, as the membership of the Romano-Latin military alliance, labelled the "Latin League" by modern scholars. But it appears that c. 500 BC there were just 15 independent Latin city-states in Latium Vetus, including Rome itself (the other 15 were annexed by the former as they expanded, especially Rome). The size of the city-state territories in c. 500 BC were estimated by Beloch (1926):
|Lavinium||Pratica di Mare, Pomezia||164|
|Fidenae||Villa Spada, Rome||51|
|Crustumerium||Marcigliana Vecchia, Settebagni||40|
|Total Size of Latium Vetus||2,347|
The table above shows the tiny size of Latium Vetus - only about two-thirds the size of the English county of Kent. Rome was by far the largest state, controlling some 35% of the total land area. The next four largest states ranged from just under half the size of Rome down to a fifth of the size; the remaining ten ranged from a tenth of the size down to less than a twentieth.
From an early stage, the external relations of the Latin city-states were dominated by their largest and most powerful member, Rome. The vast amount of archaeological evidence uncovered since the 1970s has conclusively discredited A. Alföldi's once-fashionable theory that Rome was an insignificant settlement until about 500 BC, and thus that the Republic was not established before about 450, and possibly as late as 400 BC. There is now no doubt that Rome was a unified city (as opposed to a group of separate hilltop settlements) by c. 625 BC and had become the second-largest city in Italy (after Tarentum, 510 hectares) by around 550 BC, when it had an area of about 285 hectares (1.1 sq mile) and an estimated population of 35,000. Rome was thus about half the size of contemporary Athens (585 hectares, including Piraeus) and far larger than any other Latin city.
The size of Rome at this time lends credence to the Roman tradition, dismissed by Alföldi, that in the late regal period (550–500 BC), traditionally the rule of the Tarquin dynasty, Rome established its political hegemony over the other city-states of Old Latium. According to Livy, king Tarquin the Proud bound the Latin city-states into a military alliance under Roman leadership. Reportedly, Tarquin also annexed Pometia (later Satricum) and Gabii; established control over Tusculum by a marriage alliance with its leader, Octavus Mamilius; and established Roman colonies at Signia and Circeii. He was engaged in besieging Ardea when the revolt against his monarchy broke out. Rome's political control over Latium Vetus is apparently confirmed by the text of the first recorded Romano-Carthaginian treaty, dated by the ancient Greek historian Polybius to 507 BC, a date accepted by Cornell (although some scholars argue a much later date). The treaty describes the Latin cities of Lavinium and Ardea, among others, as "Roman subjects". Although the text acknowledged that not all the Latin cities were subjects of Rome, it clearly placed them under Rome's hegemony, as it provided that if Carthage captured any Latin city, it was obliged to hand it over to Rome's control. Rome's sphere of influence is implied as extending as far as Terracina, 100 km to the south.
The fall of the Roman monarchy was probably a more lengthy, violent and international process than the swift, bloodless and internal coup related by tradition. The role in the revolution played by the Etruscan Lars Porsenna, king of Clusium, who led an invasion of Roman territory at the time of the revolution, was probably distorted for propaganda reasons by later Roman chroniclers. Livy claims that Porsenna aimed to restore Tarquin to his throne, but failed to take Rome after a siege. Tacitus suggests that Porsenna's army succeeded in occupying the City. The fact that there is no evidence of Tarquin's restoration during this occupation has led some scholars to suggest that it was Porsenna who was the real agent in the expulsion of Tarquin, and that he aimed to replace him as king of Rome. Any danger of an Etruscan takeover of Rome was removed by Porsenna's defeat at Aricia in 504 BC.
There followed a war between Rome and the other Latin city-states, which probably took advantage of the political turmoil in Rome to attempt to regain/preserve their independence. It appears that Tusculum and Aricia took the lead in organising an anti-Roman alliance. One ancient source names Egerius Baebius, the leader of Tusculum, as the "Latin dictator" (i.e. commander-in-chief of the Latin forces). It appears that Baebius dedicated a sacred grove to Diana at lucus Ferentinae (a wood near Aricia) in c. 500 BC in the presence of representatives of Latin states, including Tusculum, Aricia, Lanuvium, Lavinium, Cora, Tibur, Pometia and Ardea. This event was probably contemporaneous with, and connected with, the launch of the Latin alliance. The Latins could apparently count on the support of the Volsci Italic tribe. In addition, they were joined by the deposed Roman king Tarquin the Proud and his remaining followers.
Instead of restoring their previous hegemony, the Romans apparently settled for a military alliance on equal terms with the Latins. According to the sources, the foedus Cassianum was a bilateral treaty between the Romans on one side and the other Latin city-states combined. It provided for a perpetual peace between the two parties; a defensive alliance by which the parties pledged mutual assistance in case of attack; a promise not to aid or allow passage to each other's enemies; the equal division of spoils of war (half to Rome, half to the other Latins) and provisions to regulate trade between the parties. In addition the treaty probably provided for overall command of the allies' joint forces to alternate between a Roman and a commander from one of the other Latin city-states each year. As the nature of the Tarquinian hegemony over the Latins is unknown, it is impossible to tell how the terms of the Cassian treaty differed from those imposed by the Tarquins. But it is likely that Tarquin rule was more onerous, involving the payment of tribute, while the Republican terms simply involved a military alliance. The impetus to form such an alliance was probably provided by the acute insecurity caused by a phase of migration and invasion of the lowland areas by Italic mountain tribes in the period after 500 BC. The Latins faced repeated incursions by the Hernici, Aequi and Volsci, whose territories surrounded Latium Vetus on its eastern and southern sides.
The new Romano-Latin military alliance proved strong enough to repel the incursions of the Italic mountain tribes in the period 500–400 BC. During the succeeding century, after Rome had recovered from the catastrophic Gallic invasion of 390 BC, the Romans began a phase of expansionism. In addition to the establishment of a series of Roman colonies on territories annexed from the mountain tribes, Rome annexed a number of neighbouring Latin city-states in steady succession. The increasing threat posed by Roman encroachment led the more powerful Latin states, such as Praeneste, to attempt to defend their independence and territorial integrity by challenging Rome, often in alliance with their erstwhile enemies, mountain-tribes such as the Volsci. Finally, in 341 BC, all the Latin city-states combined in what proved to be a final effort to regain/preserve their independence. The so-called Latin War ended in 338 with a decisive Roman victory, following which Rome annexed most of Latium Vetus. A few of the larger Latin states, such as Praeneste and Tibur, were allowed to retain a degree of political autonomy, but only in a subordinate status as Roman socii ("allies"), tied to Rome by treaties of military alliance.
A genetic study published in Science in November 2019 examined the remains of six Latin males buried near Rome between 900 BC and 200 BC. They carried the paternal haplogroups R-M269, T-L208, R-311, R-PF7589 and R-P312 (two samples), and the maternal haplogroups H1aj1a, T2c1f, H2a, U4a1a, H11a and H10. These examined individuals were distinguished from preceding populations of Italy by the presence of 35% steppe ancestry. Two out of six individuals from Latin burials were found have a mixture of local Iron Age ancestry and ancestry from an Eastern mediterranean population. Among modern populations, four out of six were closest to Northern and Central Italians, and then Spaniards, while the other two were closest to Southern Italians. Overall, the genetic differentiation between the Latins, Etruscans and the preceding proto-Villanovan population of Italy was found to be insignificant.
Examined individuals from the city of Rome during the time of the Roman Empire (27 BCE – 300 CE) bore almost no genetic resemblance to Rome's founding populations, and were instead shifted towards the Eastern Mediterranean and Middle East. The Imperial population of Rome was found to have been extremely diverse, with barely any of the examined individuals being of primarily European ancestry. It was suggested that the observed genetic replacement of the city's founding populations was a result of heavy migration of merchants and slaves from the populous urban centres of the Middle East and Greece. During late antiquity, Rome's population was drastically reduced as a result of political instability, epidemics and economic changes. Repeated invasions of barbarians brought European ancestry back into Rome, resulting in the loss of genetic link to the Eastern Mediterranean and Middle East. By the Middle Ages, the people of Rome again genetically resembled European populations.
Interestingly, although Iron Age individuals were sampled from both Etruscan (n=3) and Latin (n=6) contexts, we did not detect any significant differences between the two groups with f4 statistics in the form of f4(RMPR_Etruscan, RMPR_Latin; test population, Onge), suggesting shared origins or extensive genetic exchange between them.