Neritidae Knowpia

Neritidae
	An oblique left side view of a live Theodoxus fluviatilis
	Two shells of the freshwater nerite Theodoxus danubialis, scale bar in mm
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Neritimorpha
Order:	Cycloneritida
Superfamily:	Neritoidea
Family:	Neritidae; Rafinesque, 1815
Genera
	See text
Diversity
	About 110 freshwater species, some brackish water species, and some fully marine species

Neritidae, common name the nerites, is a taxonomic family of small to medium-sized saltwater and freshwater snails which have a gill and a distinctive operculum.^[2] The family Neritidae includes marine genera such as Nerita, marine and freshwater genera such as Neritina, and freshwater and brackish water genera such as Theodoxus.

The common name "nerite" as well as the family name Neritidae and the genus name Nerita, are derived from the name of Nerites, who was a sea god in Greek mythology.

Distribution edit

Neritidae live primarily in the southern hemisphere, but there are some exceptions, such as a genus Theodoxus which can be found in Europe and Northern Africa ^[3] or Bathynerita naticoidea.

Taxonomy edit

This family consists of the five following subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):

Neritinae Rafinesque, 1815 - synonyms: Neritellinae Gray, 1847; Proto neritidae Kittl, 1899
† Neritariinae Wenz, 1938
Neritininae Poey, 1852
- tribe Neritinini Poey, 1852 - synonyms: Catilinae Gray, 1868; Orthopomatini Gray, 1868; Stenopomatini Gray, 1868; Septariini Jousseaume, 1894
- tribe Theodoxini Bandel, 2001^[4]
Smaragdiinae H. B. Baker, 1923
† Velatinae Bandel, 2001^[4]

Genera edit

Red racer nerites, Vittina waigiensis, from the Philippines. These are popular in the aquarium trade for their extremely variable shell patterns

The zebra nerite, Vittina natalensis, from East Africa. A common nerite snail in the aquarium trade and one of several species known as "zebra nerites"

Two shells of Neritodryas cornea

Genera, subgenera and species within the family Neritidae include:^[5]

† Bajanerita Squires, 1993
† Calyptronerita Le Renard, 1980
† Cuisenerita Symonds & Pacaud, 2010
† Mesoneritina Yen, 1946
† Monsneritina Kowalke, 2002

Subfamily Neritinae

Genus Bathynerita Clarke, 1989
Genus Mienerita Dekker, 2000
Genus Nerita Linnaeus, 1758 - type genus, synonyms: Turrita Wagner, 1897^[6]
- Subgenus Adenerita
- Subgenus Amphinerita
- Subgenus Cymostyla
- Subgenus Heminerita
- Subgenus Ilynerita
- Subgenus Linnerita
- Subgenus Melanerita Martens, 1889
  - Species Nerita atramentosa Reeve, 1855
  - Species Nerita melanotragus Smith, E. A., 1884
- Subgenus Nerita
- Subgenus Ritena
- Subgenus Theliostyla

Subfamily † Neritariinae

† Neritaria Koken, 1892

Subfamily Neritininae

Clithon Montfort, 1810^[7]
Clypeolum Récluz, 1842^[8]
Fluvinerita Pilsbry, 1932^[4]
- Fluvinerita tenebricosa
† Manana Gabuniya, 1949
† Mananopsis Gabuniya, 1949
Nereina de Cristofori & Jan, 1832
Neripteron Lesson, 1831^[9]
- Neripteron vespertinum (Sowerby II, 1849)

Neritina Rafinesque, 1815
- Subgenus Dostia
† Neritinopsis Gabuniya, 1953
Neritodryas Martens, 1869
Neritona Martens, 1869
† Neritoplica Oppenheim, 1892
Puperita Gray, 1857^[10]
Septaria Férussac, 1807^[11]
Theodoxus Montfort, 1810
† Velatella Meek, 1873
Vitta Mörch, 1852^[12]
Vittina H. B. Baker, 1923

Subfamily Smaragdiinae

Genus Gaillardotia Bourguignat, 1877
Genus Magadis Melvill & Standen, 1899
Genus Smaragdella Baker, 1923
Genus Smaragdia Issel, 1869^[13]
Genus Smaragoista

Subfamily † Velatinae

† Velates Montfort, 1810

Synonyms edit

† Agapilia Harzhauser & Kowalke, 2001 : synonym of Vitta Mörch, 1852
Navicella Lamarck, 1816: synonym of Septaria Férussac, 1807 (junior synonym)
† Ninniopsis Tomlin, 1930: synonym of † Theodoxus (Ninniopsis) Tomlin, 1930 represented as Theodoxus Montfort, 1810 (original rank)
Orthopoma Gray, 1868: synonym of Septaria Férussac, 1807
Orthopomatini Gray, 1868: synonym of Neritininae Poey, 1852
Paranerita Bourne, 1908: synonym of Vittina H. B. Baker, 1923 (Invalid: junior homonym of Paranerita Hampson, 1901 [Lepidoptera])
Pseudonerita H. B. Baker, 1923: synonym of Neripteron (Pseudonerita) H. B. Baker, 1923 represented as Neripteron Lesson, 1831
Serenia Benson, 1856: synonym of Neritona Martens, 1869 (unavailable name: established in synonymy)
Tanalia Gray, 1847: synonym of Neritona Martens, 1869
Theodoxia Bourguignat, 1877: synonym of Theodoxus Montfort, 1810 (Invalid: unjustified emendation of Theodoxus)
Theodoxis Montfort, 1810: synonym of Theodoxus Montfort, 1810 (alternative original spelling, not in use)

References edit

^ Strong E. E., Gargominy O., Ponder W. F. & Bouchet P. (2008). "Global Diversity of Gastropods (Gastropoda; Mollusca) in Freshwater". Hydrobiologia 595: 149-166. hdl:10088/7390 doi:10.1007/s10750-007-9012-6.
^ MolluscaBase eds. (2021). MolluscaBase. Neritininae Poey, 1852. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=411639 on 2021-09-25
^ Bunje P. M. & Lindberg D. R. (2007). "Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development". Molecular Phylogenetics and Evolution 42(2): 373–387. doi:10.1016/j.ympev.2006.06.026.
^ ^a ^b ^c Bandel K. (2001). "The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda)". Mittelungen aus dem Geologisch-Palaontologischen Institut Universitat Hamburg 85: 65-164. abstract Archived 2011-07-26 at the Wayback Machine.
^ Powell A. W. B., New Zealand Mollusca, William Collins Publishers Ltd, Auckland, New Zealand 1979 ISBN 0-00-216906-1
^ Bouchet, P.; Krijnen, C.; Gofas, S. (2016). Nerita Linnaeus, 1758. In: MolluscaBase (2017). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=138254 on 2017-12-07
^ Bouchet, P.; Rosenberg, G. (2016). Clithon Montfort, 1810. In: MolluscaBase (2017). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=737507 on 2017-12-07
^ Bouchet, P. (2016). Clypeolum. In: MolluscaBase (2017). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=873719 on 2017-12-07
^ Bouchet, P. (2016). Neripteron Lesson, 1831. In: MolluscaBase (2017). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=818409 on 2017-12-07
^ WoRMS (2009). Puperita Gray, 1857. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=205488 on 2010-05-03
^ WoRMS (2009). Septaria. Accessed through the World Register of Marine Species at http://marinespecies.org/aphia.php?p=taxdetails&id=206410 on 2010-05-03
^ Neubauer, Thomas A.; Bouchet, P.; Sartori, André F. (2016). Vitta Mörch, 1852. In: MolluscaBase (2017). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=818782 on 2017-12-07
^ WoRMS (2009). Smaragdia Issel, 1869. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2009) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=138255 on 2010-05-03

Starobogatov, Y. I. (1970). Fauna Molliuskov i Zoogeograficheskoe Raionirovanie Kontinental'nykh Vodoemov Zemnogo Shara [The Molluscan Fauna and Zoogeographical Zoning of the Continental Water Bodies of the World]. Nauka. Leningrad. 372 p., 12 tables

External links edit

Gray, J. E. (1868). Notes on Catillus, Humphrey, or Navicella, Lamarck, with descriptions of two new genera. Proceedings of the Zoological Society of London. 1867: 993-1000
Bouchet, P. & Rocroi, J.-P. (2005). Classification and nomenclator of gastropod families. Malacologia. 47 (1-2): 1-397
Bourne G. C. (1908). "Contributions to the morphology of the group Neritacea of aspidobranch gastropods. I. The Neritidae." Proceedings of the Zoological Society of London 1908: 810-887.
Miocene Gastropods and Biostratigraphy of the Kern River Area, California; United States Geological Survey Professional Paper 642 This article incorporates text from this source, which is in the public domain.

[Strong_2008-1] Strong E. E., Gargominy O., Ponder W. F. & Bouchet P. (2008). "Global Diversity of Gastropods (Gastropoda; Mollusca) in Freshwater". Hydrobiologia 595: 149-166. hdl:10088/7390 doi:10.1007/s10750-007-9012-6.

[WoRMS-2] MolluscaBase eds. (2021). MolluscaBase. Neritininae Poey, 1852. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=411639 on 2021-09-25

[Bunje_2007-3] Bunje P. M. & Lindberg D. R. (2007). "Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development". Molecular Phylogenetics and Evolution 42(2): 373–387. doi:10.1016/j.ympev.2006.06.026.

[Bandel_2001-4] Bandel K. (2001). "The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda)". Mittelungen aus dem Geologisch-Palaontologischen Institut Universitat Hamburg 85: 65-164. abstract Archived 2011-07-26 at the Wayback Machine.

[5] Powell A. W. B., New Zealand Mollusca, William Collins Publishers Ltd, Auckland, New Zealand 1979 ISBN 0-00-216906-1

[6] Bouchet, P.; Krijnen, C.; Gofas, S. (2016). Nerita Linnaeus, 1758. In: MolluscaBase (2017). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=138254 on 2017-12-07

[7] Bouchet, P.; Rosenberg, G. (2016). Clithon Montfort, 1810. In: MolluscaBase (2017). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=737507 on 2017-12-07

[8] Bouchet, P. (2016). Clypeolum. In: MolluscaBase (2017). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=873719 on 2017-12-07

[9] Bouchet, P. (2016). Neripteron Lesson, 1831. In: MolluscaBase (2017). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=818409 on 2017-12-07

[10] WoRMS (2009). Puperita Gray, 1857. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=205488 on 2010-05-03

[11] WoRMS (2009). Septaria. Accessed through the World Register of Marine Species at http://marinespecies.org/aphia.php?p=taxdetails&id=206410 on 2010-05-03

[12] Neubauer, Thomas A.; Bouchet, P.; Sartori, André F. (2016). Vitta Mörch, 1852. In: MolluscaBase (2017). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=818782 on 2017-12-07

[13] WoRMS (2009). Smaragdia Issel, 1869. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2009) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=138255 on 2010-05-03

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