Summary
Pacullidae is a family of araneomorph spiders first described by Eugène Simon in 1894.[1] It was merged into Tetrablemmidae in 1958,[2][3] then raised back to family status after a large phylogenetic study in 2017.[4]
| Pacullidae Temporal range:
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Scientific classification 
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| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Subphylum: | Chelicerata |
| Class: | Arachnida |
| Order: | Araneae |
| Infraorder: | Araneomorphae |
| Family: | Pacullidae Simon, 1894 |
| Diversity | |
| 4 genera, 39 species | |
Description edit
The family Pacullidae contains three-clawed spiders with six eyes, lacking a cribellum. They resemble spiders from the family Tetrablemmidae in some respects but are much larger, always exceeding 5 mm long, have a very wrinkled (rugose) cuticle, and females do not have large membranous receptacles.[4]
Phylogeny edit
Pacullidae falls within the Synspermiata clade, a clade of former haplogyne spiders with "synsperm" – encapsulated groups of 2–4 fused sperm cells. Within this clade, it groups with four other families, including Tetrablemmidae, but is distinct from the latter, being most closely related to Diguetidae. Together with Pholcidae, these four families are placed in the "lost trachea clade", a group of families that have lost their posterior respiratory system.[4]
| "Lost trachea clade" |
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Genera and species edit
As of April 2019[update], the World Spider Catalog accepts the following genera and species:[5]
- Lamania Lehtinen, 1981
- Lamania bernhardi (Deeleman-Reinhold, 1980) — Borneo
- Lamania bokor Schwendinger & Košulič, 2015 — Cambodia
- Lamania gracilis Schwendinger, 1989 — Bali
- Lamania inornata (Deeleman-Reinhold, 1980) — Borneo
- Lamania kraui (Shear, 1978) — Thailand, Malaysia
- Lamania lipsae Dierkens, 2011 — Borneo
- Lamania nirmala Lehtinen, 1981 — Borneo
- Lamania sheari (Brignoli, 1980) — Indonesia (Sulawesi)
- Paculla Simon, 1887
- Paculla bukittimahensis Lin & Li, 2017 — Singapore
- Paculla cameronensis Shear, 1978 — Malaysia
- Paculla globosa Lin & Li, 2017 — Singapore
- Paculla granulosa (Thorell, 1881) — New Guinea
- Paculla mului Bourne, 1981 — Borneo
- Paculla negara Shear, 1978 — Malaysia
- Paculla sulaimani Lehtinen, 1981 — Malaysia
- Paculla wanlessi Bourne, 1981 — Borneo
- Perania Thorell, 1890
- Perania annam Schwendinger & Košulič, 2015 — Vietnam
- Perania armata (Thorell, 1890) — Indonesia (Sumatra)
- Perania birmanica (Thorell, 1898) — Myanmar
- Perania cerastes Schwendinger, 1994 — Malaysia
- Perania coryne Schwendinger, 1994 — Malaysia
- Perania deelemanae Schwendinger, 2013 — Indonesia (Sumatra)
- Perania egregia Schwendinger, 2013 — Thailand
- Perania ferox Schwendinger, 2013 — Thailand
- Perania harau Schwendinger, 2013 — Indonesia (Sumatra)
- Perania korinchica Hogg, 1919 — Indonesia (Sumatra)
- Perania nasicornis Schwendinger, 1994 — Thailand
- Perania nasuta Schwendinger, 1989 — Thailand
- Perania nigra (Thorell, 1890) — Indonesia (Sumatra)
- Perania picea (Thorell, 1890) — Indonesia (Sumatra)
- Perania quadrifurcata Schwendinger, 2013 — Thailand
- Perania robusta Schwendinger, 1989 — China, Thailand
- Perania selatan Schwendinger, 2013 — Indonesia (Sumatra)
- Perania siamensis Schwendinger, 1994 — Thailand
- Perania tumida Schwendinger, 2013 — Thailand
- Perania utara Schwendinger, 2013 — Indonesia (Sumatra)
- Sabahya Deeleman-Reinhold, 1980
- Sabahya bispinosa Deeleman-Reinhold, 1980 — Borneo
- Sabahya kinabaluana Deeleman-Reinhold, 1980 — Borneo
- †Furcembolus Wunderlich 2008 9 spp, Burmese amber, Myanmar, Cenomanian
References edit
- ^ Simon, E. (1894). Histoire naturelle des araignées.
- ^ Bonnet, P. (1958). "Bibliographia araneorum". Douladoure, Toulouse. 2 (4): 3297.
- ^ Brignoli, P. M. (1973). "Ragni della Melanesia, I. Un nuovo Tetrablemma di Guadalcanal (Isole Salomone) (Araneae Tetrablemmidae)". Memorie della Società Entomologica Italiana, Genova. 72: 86.
- ^ a b c Wheeler, Ward C.; et al. (2016), "The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling", Cladistics, 33 (6): 574–616, doi:10.1111/cla.12182, PMID 34724759, S2CID 35535038
- ^ "Family: Pacullidae Simon, 1894". World Spider Catalog. Natural History Museum Bern. Retrieved 2019-04-22.