Temporal range: Late Jurassic - Early Cretaceous, 150–130 Ma
Stokesosaurus by Tom Parker.png
Artist's reconstruction of Stokesosaurus clevelandi
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Theropoda
Superfamily: Tyrannosauroidea
Clade: Pantyrannosauria
Family: Stokesosauridae
Yun & Carr, 2020
Type species
Stokesosaurus clevelandi
Madsen, 1974

Stokesosauridae is a suggested family of small to medium-sized tyrannosauroid theropods whose fossil remains have been found in North America and Europe. It includes the genera Stokesosaurus, and Juratyrant, and it may also include Eotyrannus.The family name first appeared in the supplemental info of Carr et al. (2017)[2] and was first used within the main text of a paper during Wu et al. (2019)'s description of Jinbeisaurus,[3] although no formal ICZN recognition was provided at the time.[4] The family was formally named shortly afterwards by Yun & Carr (2020).[1] The constituents of Stokesosauridae had been considered closely related to each other for several years prior. A sister-taxon relationship between Stokesosaurus and Juratyrant has been found by every major tyrannosauroid study ever since the latter was described as a species of the former in 2008.[5] Eotyrannus has also been associated with these taxa in many tyrannosauroid studies, the first of which being Brusatte & Carr (2016).[6] In addition there is a possibility which the genera Aviatyrannis and Tanycolagreus might be members of this family as well.[1]


  1. ^ a b c d Yun, Chan-Gyu; Carr, Thomas D. (2020). "Stokesosauridae clade nov., a new family name for a branch of basal tyrannosauroids". Zootaxa. 4755 (1): 195–196. doi:10.11646/zootaxa.4755.1.13.
  2. ^ Carr, Thomas D.; Varricchio, David J.; Sedlmayr, Jayc C.; Roberts, Eric M.; Moore, Jason R. (2017-03-30). "A new tyrannosaur with evidence for anagenesis and crocodile-like facial sensory system". Scientific Reports. 7 (1): 1–11. doi:10.1038/srep44942. ISSN 2045-2322.
  3. ^ Wu Xiao-chun; Shi Jian-Ru; Dong Li-Yang; Thomas D. Carr; Yi Jian; Xu Shi-Chao (2019). "A new tyrannosauroid from the Upper Cretaceous of Shanxi, China". Cretaceous Research. in press: Article 104357. doi:10.1016/j.cretres.2019.104357.
  4. ^ Mortimer, Mickey (8 March 2020). "Tyrannosauroidea". The Theropod Database.
  5. ^ Benson, R.B.J. (2008). "New information on Stokesosaurus, a tyrannosauroid (Dinosauria: Theropoda) from North America and the United Kingdom". Journal of Vertebrate Paleontology. 28 (3): 732–750. doi:10.1671/0272-4634(2008)28[732:NIOSAT]2.0.CO;2.
  6. ^ Brusatte, Stephen L.; Carr, Thomas D. (2016-02-02). "The phylogeny and evolutionary history of tyrannosauroid dinosaurs". Scientific Reports. 6 (1): 1–8. doi:10.1038/srep20252. ISSN 2045-2322.