Urocoptidae Knowpia

Urocoptidae
	Apertural view of a shell of Holospira elizabethae
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Heterobranchia
Order:	Stylommatophora
Superfamily:	Urocoptoidea
Family:	Urocoptidae; Pilsbry, 1898 (1868)

Urocoptidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Urocoptoidea.^[2]

Taxonomy edit

2005 taxonomy edit

The family Urocoptidae was classified in the superfamily Orthalicoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).^[3]

The family Urocoptidae consists of 7 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):

Urocoptinae Pilsbry, 1898 (1868) - synonym: Cylindrellidae Tryon, 1868
Apomatinae Paul, 1982
Brachypodellinae H. B. Baker, 1956
Eucalodiinae P. Fischer & Crosse, 1873
Holospirinae Pilsbry, 1946
Microceraminae Pilsbry, 1904 - synonyms: Johaniceraminae Jaume & de la Torre, 1972; Macroceraminae Jaume & de la Torre, 1972
Tetrentodontinae Bartsch, 1943

2008 taxonomy edit

Uit de Weerd (2008)^[2] moved the Urocoptidae to the newly established superfamily Urocoptoidea based on molecular phylogeny research.^[2]

The following cladogram based on Bayesian analysis of sequences of 28S ribosomal RNA shows the phylogenic relations within the Urocoptoidea.^[2] There are up to 2008 genetically analyzed species of Urocoptidae and one species of Cerionidae marked in khaki color in the cladogram.^[2]

Urocoptoidea

	Macroceramus microdon

	Microceramus pontificus

	Torrecoptis holguinensis

	Autocoptis menkeana

	Simplicervix inornata

	Brachypodella dominicensis gabbi

	Dissotropis sp. nov. [1]

	Anisospira lebmanni

Archegocoptis crenata

Eucalodium sp.

Cerionidae

Cerion striatellum

	Coelostemma sp. nov. [2]

	Hendersoniella lux lux

	Holospira fortisculpta

Epirobia polygyra

2012 taxonomy edit

Fred G. Thompson established the family Epirobiidae in 2012 for five genera of snails (Epirobia, Propilsbrya, Pectinistemma, Gyrocion, Prionoloplax), that were previously placed within Urocoptidae.^[4] Data from molecular phylogeny research as well as morphology supports monophyly of Holospiridae Pilsbry, 1946 and the Eucalodiidae Crosse & Fischer, 1868.^[4]

Genera edit

Genera in the family Urocoptidae include:

Urocoptinae

Autocoptis Pilsbry, 1902^[2]
Brachypodella L. Pfeiffer, 1840 (as Cylindrella)
Urocoptis Beck, 1837 - type genus of the family Urocoptidae^[3]

Apominae

Apoma Beck, 1837 - type genus of the subfamily Apomatinae^[3]
Simplicervix Pilsbry, 1903^[2]

Brachypodellinae

Brachypodella Beck, 1837 - type genus of the subfamily Brachypodellinae^[3]

Eucalodiinae

Anisospira Strebel, 1880^[2]
Archegocoptis Pilsbry, 1903^[2]
†Condonella McLellan, 1927^[5]
Dissotropis Bartsch, 1906^[2]
Eucalodium Crosse & P. Fischer, 1868 - type genus of the subfamily Eucalodiinae^[3]

Holospirinae

Coelostemma Dall, 1895^[2]
Hendersoniella Dall, 1905^[2]
Holospira Martens, 1860 - type genus of the subfamily Holospirinae^[3]

Microceraminae

Microceramus Pilsbry & Vanatta, 1898 - type genus of the subfamily Microceraminae^[3]
Macroceramus Guilding, 1828^[2]

Tetrentodontinae

Tetrentodon Pilsbry, 1903 - type genus of the subfamily Tetrentodontinae^[3]
Torrecoptis Bartsch, 1943^[2]

Other genera include:^[6]

Allocoptis Thompson & Franz, 1976
Amphicosmia Pilsbry & Vanatta, 1898
Anoma
Arangia
Bactrocoptis
Badiofax
Bostrichocentrum
Bostrychocentrum
Callonia
Centralia
Cochlodinella
Coelocentrum
Geoscala
Gongylostoma
Gyraxis
Haplocion
Heterocoptis
Idiostemma
Liocallonia
Metastoma
Mychostoma
Nesocoptis
Nodulia
Oligostylus
Organocoptis
Pfeiffericoptis
Pineria
Pleurostemma
Poeycoptis
Pseudopineria
Pycnoptychia Pilsbry & Vanatta, 1898
Sagracoptis
Spiroceramus
Spirostemma
Strophina
Tomelasmus
Trilamellaxis
Uncinicoptis

References edit

^ Pilsbry H. A. (1898). The Nautilus 11(9): 107.
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ Uit De Weerd, D. R. (2008). "Delimitation and phylogenetics of the diverse land-snail family Urocoptidae (Gastropoda: Pulmonata) based on 28S rRNA sequence data: A reunion with Cerion". Journal of Molluscan Studies. 74 (4): 317–29. doi:10.1093/mollus/eyn023.
^ ^a ^b ^c ^d ^e ^f ^g ^h Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
^ ^a ^b Thompson F. G. (2012). "The land snail genus Epirobia and allied genera in México and Central America, with the description of a new family, the Epirobiidae (Gastropoda, Pulmonata, Urocoptoidea)". Bulletin of the Florida Museum of Natural History 51: 167-215. PDF.
^ Roth, B. (2000). "Upper Cretaceous (Campanian) land snails (Gastropoda: Stylommatophora) from Washington and California". Journal of Molluscan Studies. 66 (3): 373–381. doi:10.1093/mollus/66.3.373.
^ "Urocoptidae - MICROCERAMIDAE". Archived from the original on 2012-07-16. Retrieved 2024-01-22.

External links edit

photos
photos
blog Bram's Snail Site with photos

[1] Pilsbry H. A. (1898). The Nautilus 11(9): 107.

[Uit_de_Weerd_2008-2] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ Uit De Weerd, D. R. (2008). "Delimitation and phylogenetics of the diverse land-snail family Urocoptidae (Gastropoda: Pulmonata) based on 28S rRNA sequence data: A reunion with Cerion". Journal of Molluscan Studies. 74 (4): 317–29. doi:10.1093/mollus/eyn023.

[Bouchet_2005-3] ^ ^a ^b ^c ^d ^e ^f ^g ^h Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.

[Thompson_2012-4] Thompson F. G. (2012). "The land snail genus Epirobia and allied genera in México and Central America, with the description of a new family, the Epirobiidae (Gastropoda, Pulmonata, Urocoptoidea)". Bulletin of the Florida Museum of Natural History 51: 167-215. PDF.

[Roth2000-5] Roth, B. (2000). "Upper Cretaceous (Campanian) land snails (Gastropoda: Stylommatophora) from Washington and California". Journal of Molluscan Studies. 66 (3): 373–381. doi:10.1093/mollus/66.3.373.

[6] "Urocoptidae - MICROCERAMIDAE". Archived from the original on 2012-07-16. Retrieved 2024-01-22.

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Summary