2004 in paleontology

Summary

Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2004.

List of years in paleontology (table)
In science
2001
2002
2003
2004
2005
2006
2007
+...

Protozoa edit

New taxa edit

Name Novelty Status Authors Age Unit Location Notes Images

Paleoleishmania[2]

Gen et sp nov

Valid

Poinar & Poinar

Albian

Burmese amber

  Myanmar

Oldest record of the protozoan family Trypanosomatidae,
Type species P. proterus

 
Paleoleishmania proterus

Fungi edit

newly named edit

Name Novelty Status Authors Age Unit Location Notes Images

Appianoporites[3]

Gen et sp nov

Valid

Smith, Currah, & Stockey

Eocene

Appian Way Flora.

  Canada

Extinct bracket fungus

Quatsinoporites[3]

Gen et sp nov

Valid

Smith, Currah, & Stockey

Barremian (Cretaceous)

Longarm Formation.

  Canada

Extinct bracket fungus.

Plants edit

Newly named plants edit

Name Novelty Status Authors Age Unit Location Notes Images

Fagus langevinii[4]

Sp nov

Valid

Manchester & Dillhoff

Ypresian

Tranquille Formation

  Canada
  British Columbia

A beech tree species

 
Fagus langevinii

Liquidambar changii[5]

Sp nov.

Valid

Pigg, Ickert-Bond, & Wen

Middle Miocene

"Yakima Canyon Flora"

  USA

A species of sweetgum

 
Liquidambar changii

Neviusia dunthornei[6]

Sp nov

Valid

DeVore, Moore, Pigg, & Wehr

Ypresian

Allenby Formation.

  Canada

oldest and only extinct species of Neviusia

Arthropoda edit

newly named arachnids edit

Name Novelty Status Authors Age Unit Location Notes Images

?Anyphops cortex[7]

sp nov

jr synonym[8]

Wunderlich

Unknown

Madagascar copal

  Madagascar

jr Synonym of Garcorops jadis

Garcorops jadis[9]

sp nov

Valid

Bosselaers

Unknown

Madagascar copal

  Madagascar

Possibly extant, but copal age is not determined

Newly named insects edit

Name Novelty Status Authors Age Unit Location Notes Images

Afromyrma[10]

Gen et sp nov

valid

Dlussky, Brothers & Rasnitsyn

Turonian

  Botswana

A Myrmicin ant, type species A. petrosa

Afropone[10]

Gen et sp nov

valid

Dlussky, Brothers & Rasnitsyn

Turonian

  Botswana

A ponerin ant, two species

Albicoccus[11]

Fam, gen et sp nov

valid

Koteja

Albian - Cenomanian

Burmese amber

  Myanmar

A scale insect, monotypic with Albicoccidae & A. dimai

Burmacoccus[11]

Fam, gen et sp nov

valid

Koteja

Albian - Cenomanian

Burmese amber

  Myanmar

A scale insect, monotypic with Burmacoccidae & B. danyi

Carebara antiqua[12]

Comb nov

valid

(Mayr, 1868)

Middle Eocene

Baltic amber

  Europe

Fossil myrmicine ant

 
Carebara antiqua

Electrostephanus janzeni[13]

Sp nov

jr synonym

Engel

Lutetian

Baltic Amber

  Russia

A stephanid wasp

Gerontoformica[14]

Gen et sp nov

valid

Nel & Perrault

Albian - Cenomanian

Charentese amber

  France

A stem group ant,
type species G. cretacica

 
Gerontoformica cretacica

Marmyan[11]

gen et sp nov

valid

Koteja

Cenomanian

Burmese amber

  Myanmar

A scale insect, monotypic M. barbarae

Microberotha[15]

Gen et sp nov

Valid

Archibald

Ypresian

Hat Creek Amber

  Canada

A Berothid lacewing

Palaeomyia[16]

Gen et sp nov

Valid

Poinar

Cenomanian

Burmese amber

  Myanmar

A Phlebotomidae sand fly, type species P. burmitis

 
Palaeomyia burmitis

Newly named fishes edit

Name Novelty Status Authors Age Unit Location Notes Images

Gomphonchus? minutus[17]

Sp. nov

Valid

Valiukevičius

Late Silurian

Minija Formation

  Latvia

An acanthodian belonging to the group Ischnacanthiformes and the family Ischnacanthidae.

Lophosteus canadensis[18]

Sp. nov

Valid

Schultze & Märss

Late Silurian

Barlow Inlet Formation

  Canada

A bony fish belonging to the group Lophosteiformes.

Lophosteus ohesaarensis[18]

Sp. nov

Valid

Schultze & Märss

Late Silurian

  Estonia
  Latvia

A bony fish belonging to the group Lophosteiformes.

Lophosteus uralensis[18]

Sp. nov

Valid

Schultze & Märss

Early Devonian

  Russia

A bony fish belonging to the group Lophosteiformes.

Nostolepis alifera[17]

Sp. nov

Valid

Valiukevičius

Late Silurian

Targale Formation

  Latvia

An acanthodian belonging to the group Climatiiformes and the family Climatiidae.

Nostolepis latvica[17]

Sp. nov

Valid

Valiukevičius

Late Silurian

Minija Formation
Targale Formation

  Latvia

An acanthodian belonging to the group Climatiiformes and the family Climatiidae.

Perscheia[19]

Gen. et sp. nov

Valid

Elliott, Mark-Kurik & Daeschler

Devonian (Frasnian)

Nordstrand Point Formation

  Canada

A member of Pteraspidiformes belonging to the group Psammosteida and the family Obrucheviidae. The type species is P. pulla

Poracanthodes marssae[17]

Sp. nov

Valid

Valiukevičius

Late Silurian

Targale Formation

  Estonia
  Latvia

An acanthodian belonging to the group Ischnacanthiformes and the family Poracanthidae.

Poracanthodes sulcatus[17]

Sp. nov

Valid

Valiukevičius

Late Silurian

Targale Formation

  Estonia
  Latvia

An acanthodian belonging to the group Ischnacanthiformes and the family Poracanthidae.

Talimaalepis[20]

Gen. et sp. nov

Valid

Žigaitė

Silurian

Dashtygoi Formation
Kyzyl-Tchiraa Formation
Tchergak Formation

  Mongolia
  Russia

A thelodont, possibly a member of the group Phlebolepidiformes. The type species is T. rimae

Trimpleylepis[21]

Gen. et 2 sp. nov

Valid

Miller, Märss & Blom

Late Silurian

  Estonia
  United Kingdom

A member of Anaspida belonging to the group Birkeniida, possibly a member of the family Septentrioniidae. The type species is T. juncta; genus also includes T. concatenata.

Newly named amphibians edit

Name Novelty Status Authors Age Unit Location Notes Images

Jakubsonia[22]

Gen. et sp. nov

Valid

Lebedev

Devonian (Famennian)

  Russia

A basal member of Tetrapoda. The type species is J. livnensis

Reptile edit

Archosauromorphs edit

Newly named dinosaurs edit

Data courtesy of George Olshevsky's dinosaur genera list.[23]

Name Novelty Status Authors Age Unit Location Notes Images

Atrociraptor[24]

Valid

Currie & Varricchio

Edmontonian

Horseshoe Canyon Formation

  Canada

 
Atrociraptor

Bashunosaurus[25]

Gen et sp nov

Valid

Kuang

Middle Jurassic

Xiashaximiao Formation

  China

Bissektipelta[26]

Gen et sp nov

Valid

Parish & Barrett

Cenomanian-Turonian

Bissekty Formation

  Uzbekistan

Bonatitan[27]

Gen et sp nov

Valid

Martinelli & Forasiepi

Late Cretaceous

Bajo de Santa Rosa

  Argentina

Bonitasaura[28]

Gen et sp nov

Valid

Apesteguía

late Cretaceous

Bajo de la Carpa Formation

  Argentina

 
Bonitasaura

Borealosaurus[29]

Gen et sp nov

Valid

You, Ji, Lamanna,
Li, & Li

late Cretaceous

Sunjiawan Formation

  China

Dilong[30]

Gen et sp nov

Valid

Xu, Norell, Kuang,
Wang, Zhao, & Jia

Early Cretaceous

Yixian Formation

  China

 
Dilong

Ekrixinatosaurus[31]

Gen et sp nov

Valid

Calvo, Rubilar-Rogers, & Moreno

Late Cretaceous

Candeleros Formation

  Argentina

 
Ekrixinatosaurus

Graciliraptor[32]

Valid

Xing & Wang

Barremian

Yixian Formation

  China

Huaxiagnathus[33]

Valid

Hwang, Norell, Ji, & Gao

Barremian

Yixian Formation

  China

 
Huaxiagnathus

Kerberosaurus[34]

Valid

Bolotsky & Godefroit

Late Cretaceous

  Russia

Limaysaurus[35]

Valid

Salgado, Garrido, Cocca, & Cocca

  Argentina

 
Limaysaurus

Mei[36]

Valid

Xu & Norell

Barremian

Yixian Formation

  China

 
Mei

Mirischia[37]

Valid

Naish, Martill, & Frey

Albian

Santana Formation

  Brazil

 
Mirischia

Nemegtia[38]

Preoccupied name

Lü, Tomida, Azuma,
Dong, Lee, & Szczechura

Renamed Nemegtomaia

Otogosaurus

Nomen nudum

Zhao & Tan

Prenoceratops[39]

Valid

Chinnery

  USA
  Montana

 
Prenoceratops

Rugops[40]

Valid

Sereno, Wilson, & Conrad

  Niger

 
Rugops

Sinusonasus[41]

Valid

Xu, & Wang

Barremian

Yixian Formation

  China

Spinostropheus[40]

Valid

Sereno, Wilson & Conrad

Middle Jurassic

Irhazer Group

  Niger

A ceratosaur; new genus for
"Elaphrosaurus" gautieri Lapparent 1960.

Suuwassea[42]

Valid

Harris & Dodson

Kimmeridgian

Morrison Formation

  United States

 
Suuwassea

Talenkauen[43]

Valid

Novas, Cambiaso, & Ambrosio

Pari Aike Formation

  Argentina

 
Talenkauen

Tazoudasaurus[44]

Valid

Allain et al

  Morocco

Unaysaurus[45]

Valid

Leal, Azevedo,
Kellner, & Da Rosa

Caturrita Formation

  Brazil

 
Unaysaurus

Pterosaurs edit

New taxa edit
Name Status Authors Age Unit Location Notes

Avgodectes

Valid

Peters

Early Cretaceous

Yixian Formation

  China

A pterosaur hatchling.

Cacibupteryx

Valid

Gasparini Fernández de la Fuente

Late Jurassic (Oxfordian)

Jagua Formation

  Cuba

A rhamphorhynchid.

"Daitingopterus"

Nomen Nudum

Maisch Matzke Ge Sun

Late Jurassic (Tithonian)

Solnhofen Limestone

  Germany

A germanodactylid.

Lonchognathosaurus

Valid

Maisch, M.W., Matzke, A.T., and Ge Sun

Early Cretaceous (Aptian)

Lianmuqin Formation

  China

A dsungaripterid.

Other archosauromorphs edit

Name Status Authors Location Notes Images
Crosbysaurus[46] Valid non-dinosaurian taxon.
  • Heckert
  USA
Protecovasaurus[47] Valid non-dinosaurian taxon.
  • Heckert
  USA

Newly named birds edit

Name Status Novelty Authors Age Unit Location Notes Images

Aberratiodontus [48]

Valid

Gen et sp. nov.

Gong, Hou & Wang

Early Cretaceous

Jiufotang Formation

  China

An Enantiornithes,
Type species A. wui

Anserpica kiliani [49]

Valid

Gen et sp. nov.

Cécile Mourer-Chauviré

Didier Berthet

Marguerite Hugueney

Late Oligocene, Early Miocene

MP 29

  France

An Anseranatidae, the type species of Anserpica Mourer-Chauviré, Berthet & Hugueney, 2004.

Bavaripsitta ballmanni [50]

Valid

Gen et sp. nov.

Gerald Mayr

Ursula B. Göhlich

Middle Miocene

Mn 6

  Germany:

  Bavaria

A Psittacidae, the type species of Bavaripsitta Mayr & Göhlich, 2004.

Borvocarbo guilloti [49]

Valid

Gen et sp. nov.

Cécile Mourer-Chauviré

Didier Berthet

Marguerite Hugueney

Late Oligocene, Early Miocene

MP 30, MN 1

  France;

  Germany

A Phalacrocoracidae, the type species of Borvocarbo Mourer-Chauviré, Berthet & Hugueney, 2004.

?Colymboides metzleri [51]

Valid

Sp. nov.

Gerald Mayr

Early Oligocene

Rupelian

  Germany:

  Baden-Württemberg;

  Belgium:

  Antwerp

A Gaviidae, not certain a species of Colymboides Milne-Edwards, 1868.

Eurotrochilus inexpectatus [52]

Valid

Gen et sp. nov.

Gerald Mayr

Early Oligocene

Rupelian

  Germany:

  Baden-Württemberg

A Trochilidae, the type species of Eurotrochilus Mayr, 2004.

Iaceornis marshi [53]

Valid

Gen et sp. nov.

Julia A. Clarke

Late Cretaceous

Niobrara Formation

  USA:

  Kansas

An Ornithurae, the type species of Iaceornis Clarke, 2004.

Longirostravis hani [54]

Valid

Gen et sp. nov.

Hou Lianhai

Luis M. Chiappe

Zhang Fucheng

Chuong Cheng-Ming

Early Cretaceous

Yixian Formation

  China

An Enantiornithes, Euenantiornithes, the type species of Longirostravis Hou, Chiappe, Zhang et Chuong, 2004.

Meridiocichla salotti [55]

Valid

Gen et sp. nov.

Antoine Louchart

Late Quaternary

Corsica, Crete

  France:

  Corsica;

  Greece

A Turdidae, the type species of Meridiocichla Louchart, 2004.

Miopica paradoxa [56]

Valid

Gen et sp. nov.

Evgeny N. Kurochkin

Denis V. Sobolev

Miocene

Middle Meothis

MN 10

  Ukraine

A Corvidae, the type species of Miopica Kurochkin & Sobolev, 2004.

Mirolia brevirostrata [57]

Valid

Gen et sp. nov.

Peter Ballmann

Middle Miocene

MN 6

  Germany:

  Bavaria

A Scolopacidae, the type species of Mirolia Ballmann, 2004.

Mirolia dubia [57]

Valid

Sp. nov.

Peter Ballmann

Middle Miocene

MN 6

  Germany:

  Bavaria

A Scolopacidae.

?Mirolia mascalidris [57]

Valid

Sp. nov.

Peter Ballmann

Middle Miocene

MN 6

  Germany:

  Bavaria

A Scolopacidae, not certain to belong in Mirolia Ballmann, 2004.

Mirolia parvula [57]

Valid

Sp. nov.

Peter Ballmann

Middle Miocene

MN 6

  Germany:

  Bavaria

A Scolopacidae.

Primobucco frugilegus [58]

Valid

Sp. nov.

Gerald Mayr

Cécile Mourer-Chauviré

Ilka Weidig

Middle Eocene

Messel pit,

MP 11

  Germany:

  Hessen

A Primobucconidae Feduccia et Martin, 1976.

Primobucco perneri [58]

Valid

Sp. nov.

Gerald Mayr

Cécile Mourer-Chauviré

Ilka Weidig

Middle Eocene

Messel pit

MP 11

  Germany:

  Hessen

A Primobucconidae Feduccia et Martin, 1976.

Ramphastosula ramirezi [59]

Valid

Gen et sp. nov.

Marcelo Stucchi

Mario Urbina

Early Pliocene

Pisco Formation

  Peru

A Sulidae, the type species of Ramphastosula Stucchi & Urbina, 2004.

Tyto mourerchauvireae [60]

Valid

Sp. nov.

Marco Pavia

Early-Middle Pleistocene

Sicily

  Italy:

  Sicily

A Tytonidae.

Vescornis hebeiensis [61]

Valid

Gen et sp. nov.

Zhang Fucheng

Per G.P. Ericson

Zhou Zhonghe

Early Cretaceous

Yixian Formation

  China

An Enentiornithes Walker, 1981, this is the type species of the new genus, most probably a junior synonym of Hebeiornis fengningensis Yan, 1999.

Vitirallus watlingi [62]

Valid

Gen et sp. nov.

Trevor H. Worthy

Holocene

Viti Levu

  Fiji

A Rallidae, the type species of Vitirallus Worthy, 2004.

Wingegyps cartellei [63]

Valid

Gen et sp. nov.

Herculano M. F. de Alvarenga

Storrs L. Olson

Late Pleistocene or Early Holocene

Gruta dos Brejões

  Brazil

A Cathartidae, the type species of Wingegyps Alvarenga & Olson, 2004.

Plesiosaurs edit

New taxa edit

Name Status Authors Age Unit Location Notes
Plesiopterys Valid O'Keefe Toarcian Posidonienschiefer   Germany

Synapsids edit

Non-mammalian edit

Name Status Authors Age Unit Location Notes Images
Ianthodon[64] Valid Kissel & Reisz Kasimovian   USA
 
Lobalopex
Lobalopex[65] Valid Sidor, Hopson and Keyser Lopingian Teekloof Formation   South Africa
Progalesaurus[66] Valid Sidor & Smith Induan Balfour Formation   South Africa
Pyozia [67] Valid Anderson and Reisz Capitanian Krasnoschelsk Formation   Russia
Rewaconodon[68] Valid Datta, Das & Luo Late Triassic Tiki Formation   India

Other Animals edit

Name Status Authors Age Unit Location Notes Images
Charniodiscus procerus Valid LaFlamme et al., 2004 Ediacaran Mistaken Point Formation and Trepassey Formation   Canada

References edit

  1. ^ Gini-Newman, Garfield; Graham, Elizabeth (2001). Echoes from the past: world history to the 16th century. Toronto: McGraw-Hill Ryerson Ltd. ISBN 9780070887398. OCLC 46769716.
  2. ^ Poinar, G. (2008). "Lutzomyia adiketis sp. n. (Diptera: Phlebotomidae), a vector of Paleoleishmania neotropicum sp. n. (Kinetoplastida: Trypanosomatidae) in Dominican amber". Parasites & Vectors. 1 (1): 22. doi:10.1186/1756-3305-1-22. PMC 2491605. PMID 18627624.
  3. ^ a b Smith, S.Y.; Currah, R.S.; Stockey, R.A. (2004). "Cretaceous and Eocene poroid hymenophores from Vancouver Island, British Columbia". Mycologia. 96 (1): 180–186. doi:10.2307/3762001. JSTOR 3762001. PMID 21148842.
  4. ^ Manchester, S. R.; Dillhoff, R. M. (2004). "Fagus (Fagaceae) fruits, foliage, and pollen from the Middle Eocene of Pacific Northwestern North America". Canadian Journal of Botany. 82 (10): 1509–1517. doi:10.1139/b04-112.
  5. ^ Pigg, K. B.; Ickert-Bond, S. M.; Wen, J. (2004). "Anatomically preserved Liquidambar (Altingiaceae) from the middle Miocene of Yakima Canyon, Washington state, USA, and its biogeographic implications". American Journal of Botany. 91 (3): 499–509. doi:10.3732/ajb.91.3.499. PMID 21653405.
  6. ^ DeVore, M.L.; Moore, S.M.; Pigg, K.B.; Wehr, W.C. (2004). "Fossil Neviusia leaves (Rosaceae: Kerrieae) from the Lower Middle Eocene of Southern British Columbia". Rhodora. 12 (927): 197–209. JSTOR 23314752.
  7. ^ Wunderlich, J. (2004). "Fossil spiders in amber and copal. Conclusions, revisions, new taxa and family diagnoses of fossil and extant taxa". Beiträge zur Araneologie. 3AB: 1–1908.
  8. ^ Penney, D.; Ono, H.; Selden, P.A. (2005). "A new synonymy for the Madagascan copal spider fauna (Araneae, Selenopidae)" (PDF). Journal of Afrotropical Zoology. 2: 41–44.[permanent dead link]
  9. ^ Bosselaers, J. (2004). "A new Garcorops species from Madagascar copal (Araneae: Selenopidae)" (PDF). Zootaxa. 445: 1–7. doi:10.11646/zootaxa.445.1.1. S2CID 47620450.
  10. ^ a b Dlussky, G. M.; Brothers, D. J.; Rasnitsyn, A. P. (2004). "The first Late Cretaceous ants (Hymenoptera: Formicidae) from southern Africa, with comments on the origin of the Myrmicinae". Insect Systematics and Evolution. 35: 1–13. doi:10.1163/187631204788964727.
  11. ^ a b c Koteja, J (2004). "Scale insects (Hemiptera: Coccinea) from cretaceous Myanmar (Burmese) amber". Journal of Systematic Palaeontology. 2 (2): 109–114. Bibcode:2004JSPal...2..109K. doi:10.1017/S1477201904001166. S2CID 86393735. – via Taylor & Francis (subscription required)
  12. ^ Fernández, F. (2004). "The American species of the myrmicine ant genus Carebara Westwood (Hymenoptera: Formicidae)". Caldasia. 26: 191–238.
  13. ^ Engel, M.S.; Ortega-Blanco, J. (2008). "The fossil crown wasp Electrostephanus petiolatus Brues in Baltic Amber (Hymenoptera, Stephanidae): designation of a neotype, revised classification, and a key to amber Stephanidae". ZooKeys (4): 55–64. Bibcode:2008ZooK....4...55E. doi:10.3897/zookeys.4.49. hdl:2445/36428.
  14. ^ Nel, A.; Perrault, G.; Perrichot, V.; Néradeau, D. (2004). "The oldest ant in the lower cretaceous amber of Charente-maritime (SW France) (Insecta: Hymenoptera: Formicidae)". Geologica Acta. 2 (1): 23–29.
  15. ^ Archibald, S.B.; Makarkin, V.N. (2004). "New genus of minute Berothidae (Neuroptera) from Early Eocene amber of British Columbia" (PDF). The Canadian Entomologist. 136: 61–76. CiteSeerX 10.1.1.552.2285. doi:10.4039/n03-043. S2CID 36459014. Archived from the original (PDF) on 2011-08-18. Retrieved 2010-04-09.
  16. ^ Poinar, G. O. Jr. (2004). "Palaeomyia burmitis (Diptera: Phlebotomidae), a new genus and species of Cretaceous sand flies with evidence of blood-sucking habits". Proceedings of the Entomological Society of Washington. 106: 598–605.
  17. ^ a b c d e Valiukevičius, Juozas (2004). "Silurian acanthodian succession of the Lužni-4 borehole (Latvia)" (PDF). Acta Universitatis Latviensis. Earth and Environment Sciences. 679: 120–147.
  18. ^ a b c Schultze, Hans-Peter; Märss, Tiiu (2004). "Revisiting Lophosteus, a primitive osteichthyan" (PDF). Acta Universitatis Latviensis. Earth and Environment Sciences. 679: 57–78.
  19. ^ Elliott, David K.; Mark-Kurik, Elga; Daeschler, Edward B. (2004). "A revision of Obruchevia (Psammosteida: Heterostraci) and a description of a new obrucheviid from the Late Devonian of the Canadian Arctic" (PDF). Acta Universitatis Latviensis. Earth and Environment Sciences. 679: 22–45.
  20. ^ Žigaitė, Živilė (2004). "A new thelodont from Lower Silurian of Tuva and north-west Mongolia" (PDF). Acta Universitatis Latviensis. Earth and Environment Sciences. 679: 158–165.
  21. ^ Miller, C. Giles; Märss, Tiiu; Blom, Henning (2004). "New anaspid material from the Late Silurian of Britain and Estonia" (PDF). Acta Universitatis Latviensis. Earth and Environment Sciences. 679: 46–56.
  22. ^ Lebedev, Oleg A. (2004). "A new tetrapod Jakubsonia livnensis from the Early Famennian (Devonian) of Russia and palaeoecological remarks on the Late Devonian tetrapod habitats" (PDF). Acta Universitatis Latviensis. Earth and Environment Sciences. 679: 79–98.
  23. ^ Olshevsky, George. "Dinogeorge's Dinosaur Genera List". Archived from the original on 2011-07-15. Retrieved 2008-08-07.
  24. ^ Currie, P.J. and D.J. Varricchio. 2004. A new dromaeosaurid from the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta, Canada. In: Feathered Dragons: Studies on the Transition from Dinosaurs to Birds (P.J. Currie, E.B. Koppelhus, M.A. Shugar, and J.L. Wright, eds.). Indiana University Press, Bloomington, IA: pp. 112-132.
  25. ^ Kuang, X.W. (2004). "A new Sauropoda from Kaijiang dinosaur fauna in middle Jurassic beds of North-Eastern Sichuan". In Sun, J.W. (ed.). Collection of the 90th anniversary of Tianjin museum of natural history. Tianjin: Tianjin Science and Technology Press. pp. 40–46.
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