With over 1700 recognized recent and fossil species,[2] this is among the most diverse families of terrestrial gastropods (cf. Orthalicidae), although the marine gastropod family Pyramidellidae is larger.
Most species of Clausiliidae have an anatomical structure known as a clausilium, which enables the snail to close off the aperture of the shell with a sliding "door".
Shell descriptionedit
Almost all the species of snails in the family of door snails are left-handed, which is an uncommon feature in gastropod shells in general.
These snails have shells which are extremely high-spired, with numerous whorls.
The shells tend to be club-shaped, tapering at both ends to a rounded nub. The aperture usually has visible folds.
The clausiliumedit
Clausiliids are also very unusual among pulmonate gastropods in that most of them have a "door" or clausilium. The clausilium is not the same thing as an operculum, which does not exist at all in pulmonate gastropods.
The clausilium is a calcareous structure, tongue-shaped or spoon-shaped, which can close the aperture of the snail shell to protect the soft parts against predation by animals such as carnivorous beetle larvae. The narrow end of the clausilium slides in the grooves that are formed by the folds on the inside of the shell.
Anatomyedit
In this family, the number of haploidchromosomes lies between 21 and 30 (according to the values in this table).[3]
Cochlodina A. Férussac, 1821 - with four subgenera: Cochlodina; Cochlodinastra H. Nordsieck, 1977; Paracochlodina H. Nordsieck, 1969; Procochlodina H. Nordsieck, 1969
Bathyptychia Lindholm, 1925 - with three subgenera: Bathyptychia; Brachyptychia H. Nordsieck, 2001 and Strictiphaedusa H. Nordsieck, 2001
Castanophaedusa Páll-Gergely & Szekeres, 2017
Celsiphaedusa H. Nordsieck, 2001
Changphaedusa Motochin & Ueshima, 2017
Cylindrophaedusa O. Boettger, 1877 - with three subgenera: Cylindrophaedusa and Montiphaedusa H. Nordsieck, 2002
Dautzenbergiella Lindholm, 1924 - with two subgenera: Dautzenbergiella and Mansuyiella H. Nordsieck, 2003
Euphaedusa O. Boettger, 1877 - with subgenera: Dentiphaedusa H. Nordsieck, 2003; Euphaedusa; and Telophaedusa H. Nordsieck, 2003
Fuchsiana Gredler, 1887
Hemiphaedusa O. Boettger, 1877 - with subgenera: Dendrophaedusa H. Nordsieck, 2002; Hemiphaedusa; Hemiphaedusoides H. Nordsieck, 2001; Hemizaptyx Pilsbry, 1905; Labyrinthiphaedusa H. Nordsieck, 2001; ; Pinguiphaedusa Azuma, 1982; Placeophaedusa Minato, 1994)
Juttingia Loosjes, 1965 - with two subgenera: Juttingia and Pseudohemiphaedusa H. Nordsieck, 2002
Liparophaedusa Lindholm, 1924
Loosjesia H. Nordsieck, 2002
Luchuphaedusa Pilsbry, 1901 - with two subgenera: Luchuphaedusa and Nesiophaedusa Pilsbry, 1905
Macrophaedusa Moellendorff, 1883
Macrophaedusella H. Nordsieck, 2001
Megalophaedusa O. Boettger, 1877 - with four subgenera: Megalophaedusa; Mesophaedusa Ehrmann, 1929; Mesozaptyx Kuroda, 1963 and Mundiphaedusa Minato, 1979. (Neophaedusa is a synonym of Megalophaedusa).[14]
Oospira Blanford, 1872[15] - with subgenera Formosanella H. Nordsieck, 2003; Oospira; Paraformosella H. Nordsieck, 2003 and Siphonophaedusa Lindholm, 1924
Papilliphaedusa H. Nordsieck, 2003
Paraphaedusa O. Boettger, 1877
Phaedusa H. & A.Adams, 1855 - with subgenera: Metaphaedusa H. Nordsieck, 2001; Phaedusa; Pseudophaedusa Tomiyama, 1984
Probosciphaedusa Z.-Y. Chen, 2021
Reinia Kobelt, 1876 - with three subgenera: Parareinia H. Nordsieck, 1998; Pictophaedusa Azuma, 1982 and Reinia
Serriphaedusa H. Nordsieck, 2001
Sinigena Lindholm, 1925
Solitariphaedusa Motochin & Ueshima, 2017
Stereophaedusa O. Boettger, 1877
Streptodera Lindholm, 1925
Tauphaedusa H. Nordsieck, 2003
Tosaphaedusa Ehrmann, 1929
Zaptyx Pilsbry, 1901 - with two subgenera: Prozaptyx Loosjes, 1950 and Zaptyx
Leptacme Ehrmann, 1927:[15] synonym of Oospira (Leptocochlea) Grego & Szekeres, 2011 represented as Oospira W. T. Blanford, 1872
Lindholmiella Ehrmann, 1927: synonym of Oospira (Lindholmiella) Ehrmann, 1927 represented as Oospira W. T. Blanford, 1872
Metazaptyx Pilsbry, 1905synonym of Zaptyx Pilsbry, 1900
Neniauchenia H. Nordsieck, 2002: synonym of Grandinenia Minato & D.-N. Chen, 1984
Neostyriaca A. J. Wagner, 1920:[9] synonym of Clausilia (Neostyriaca) A. J. Wagner, 1920 represented as Clausilia Draparnaud, 1805
Oligozaptx Pilsbry, 1905: synonym of Zaptyx (Stereozaptyx) Pilsbry, 1905 represented as Zaptyx Pilsbry, 1900
Parazaptyx Pilsbry, 1905: synonym of Zaptyx (Parazaptyx) Pilsbry, 1905 represented as Zaptyx Pilsbry, 1900
Pliciphaedusa H. Nordsieck, 1998: synonym of Stereophaedusa (Pliciphaedusa) H. Nordsieck, 1998 represented as Stereophaedusa O. Boettger, 1877
Pulchraptyx Minato, 1981: synonym of Zaptyx (Pulchraptyx) Minato, 1981 represented as Zaptyx Pilsbry, 1900
Renschiphaedusa Loosjes & Loosjes-van Bemmel, 1973: synonym of Phaedusa H. Adams & A. Adams, 1855
Selenoptyx Pilsbry, 1908: synonym of Zaptyx (Selenoptyx) Pilsbry, 1908 represented as Zaptyx Pilsbry, 1900
Serrulinella H. Nordsieck, 1984: synonym of Serrulina (Serrulinella) H. Nordsieck, 1984 represented as Serrulina Mousson, 1873 (unaccepted rank)
Stereozaptyx Pilsbry, 1905: synonym of Zaptyx (Stereozaptyx) Pilsbry, 1905 represented as Zaptyx Pilsbry, 1900
Thaumatoptyx Pilsbry, 1908: synonym of Zaptyx (Thaumatoptyx) Pilsbry, 1908 represented as Zaptyx Pilsbry, 1900
Tyrannophaedusa Pilsbry, 1901: synonym of Megalophaedusa (Tyrannophaedusa) Pilsbry, 1900 represented as Megalophaedusa O. Boettger, 1877
Tyrannozaptyx Käufel, 1930: synonym of Zaptyx (Tyrannozaptyx) Käufel, 1930 represented as Zaptyx Pilsbry, 1900
Zaptychopsis Ehrmann, 1927: synonym of Zaptyx (Zaptychopsis) Ehrmann, 1927 represented as Zaptyx Pilsbry, 1900
Conservationedit
Although non-marine molluscs appear to be exceptionally vulnerable to extinction,[16] the IUCN Red list listed only 9 species[17] from this family.
Referencesedit
^J. E. Gray. April 14, 1855. Catalogue of Pulmonata or air-breathing Mollusca in the collection of the British Museum, Part I: 156.
^ abMolluscaBase eds. (2023). MolluscaBase. Clausiliidae L. Pfeiffer & J. E. Gray, 1855. Accessed through: World Register of Marine Species at: https://www.molluscabase.org/aphia.php?p=taxdetails&id=833936 on 2023-04-11
^Barker G. M.: Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. in Barker G. M. (ed.): The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, 2001, ISBN 0-85199-318-4. 1-146, cited pages: 139 and 142.
^Rodrigo B. Salvador; Abraham S.H. Breure (2020). "Type material of Clausiliidae door snails from Philippe Dautzenberg in the collection of the Museum of New Zealand Te Papa Tongarewa". Tuhinga: Records of the Museum of New Zealand Te Papa Tongarewa. 31. Te Papa: 56–69. ISSN 1173-4337. Wikidata Q106839645.
^ abcdefghi"Clausiliini". Fauna Europaea, last update 27 January 2011, accessed 27 April 2011.
^"Graciliaria". Fauna Europaea, last update 27 January 2011, accessed 27 April 2011.
^Uit de Weerd D. R. (2004). "Molecular phylogenetic history of eastern Mediterranean Alopiinae, a group of morphologically indeterminate land snails". Doctoral thesis, Leiden University, ISBN 90-6464-874-3.
^Gittenberger E. & Uit de Weerd D. R. (2009). "Summarizing data on the Inchoatia taxa, including Inchoatia megdova bruggeni subspec. nov. (Gastropoda, Pulmonata, Clausiliidae)" Zoologische Mededelingen83 http://www.zoologischemededelingen.nl/83/nr03/a08 Archived 16 January 2014 at the Wayback Machine
^Rodrigo B. Salvador; Alina Wahab; Nicole E. Phillips; Abraham S.H. Breure (2021). "South American and Trinidadian terrestrial Gastropoda in the collection of the Museum of New Zealand Te Papa Tongarewa". Tuhinga: Records of the Museum of New Zealand Te Papa Tongarewa. 32. Te Papa: 64–80. ISSN 1173-4337. Wikidata Q116264204.
^cf. Nordsieck H. (2006). "Species list of recent Clausiliidae".
^ abMaassen W. J. M. & Gittenberger E. (2007). "Three new clausiliid land snails from Tonkin, northern Vietnam (Gastropoda: Pulmonata: Clausiliidae)". Zoologische Mededelingen81(1): http://www.zoologischemededelingen.nl/81/nr01/a10 Archived 7 October 2011 at the Wayback Machine
^Lydeard, C.; Cowie R.; Ponder, W.F.; et al. (April 2004). "The global decline of nonmarine mollusks". BioScience. 54 (4): 321–330. doi:10.1641/0006-3568(2004)054[0321:TGDONM]2.0.CO;2.
^IUCN 2009. IUCN Red List of Threatened Species. Version 2009.2. <www.iucnredlist.org>. Downloaded on 5 March 2010.
Further readingedit
Maltz T. K. & Sulikowska-Drozd A. (2008) "Life Cycles of Clausiliids of Poland — Knowns and Unknowns". Annales Zoologici58(4): 857-880. doi:10.3161/000345408X396783.
Nordsieck H. (2007). Worldwide Door Snails. ConchBooks, 213 pp. ISBN 978-3-939767-07-7.
Uit de Weerd D. R. (2004). "Molecular phylogenetic history of eastern Mediterranean Alopiinae, a group of morphologically indeterminate land snails". Doctoral thesis, Department of Biology, Faculty of Science, Leiden University. HTM, PDF.
Uit de Weerd D. R., Piel W. H. & Gittenberger E. (2004). "Widespread polyphyly among Alopiinae snail genera: when phylogeny mirrors biogeography more closely than morphology". Molecular Phylogenetics and Evolution33(3): 533-548. doi:10.1016/j.ympev.2004.07.010
Páll-Gergely B. & Szekeres M. , 2017. New and little-known Clausiliidae (Gastropoda: Pulmonata) from Laos and southern Vietnam. Journal of Conchology 42(6): 507-519
External linksedit
Wikimedia Commons has media related to Clausiliidae.